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Caypullisaurus Temporal range: Tithonian-Berriasian ~150–140.2 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Dakosaurus (right) pursuing a Caypullisaurus (left)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	†Ichthyosauria
Family:	†Ophthalmosauridae
Subfamily:	†Platypterygiinae
Genus:	†Caypullisaurus Fernández, 1997
Species:	†C. bonapartei
Binomial name
†Caypullisaurus bonapartei Fernández, 1997

Caypullisaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic to the Early Cretaceous (Tithonian and Berriasian stages) of Argentina. Its holotype was collected from the Vaca Muerta Formation of Cerro Lotena, Neuquen, dating to the early Tithonian stage of the Late Jurassic, about 150 million years ago. Caypullisaurus was first named by Marta Fernández in 1997 and the type species is Caypullisaurus bonapartei. It was a large ichthyosaur, measuring about 7 m (23 ft) long. The forelimbs of Caypullisaurus contained 10 digits each.

Classification

It is a member of the family Ophthalmosauridae, and closely related to Platypterygius and Brachypterygius. In 2012, Caypullisaurus was found to be most closely related to Athabascasaurus and "Platypterygius" australis, and to nest within the subfamily Platypterygiinae.

Phylogeny

The following cladogram shows a possible phylogenetic position of Caypullisaurus in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).

See also

• List of ichthyosaurs
• Timeline of ichthyosaur research

References

1. Fernández, M.S. (1997). "A new Ichthyosaur from the Tithonian (Late Jurassic) of the Neuquen Basin, Northwestern Patagonia, Argentina". Journal of Paleontology. 71 (3): 479–484. Bibcode:1997JPal...71..479F. doi:10.1017/S0022336000039494.
2. Gasparini, Z.; Fernández, M. (2005). "Jurassic marine reptiles of the Neuquén Basin: records, faunas and their palaeobiogeographic significance". Geological Society, London, Special Publications. 252 (1): 279–294. Bibcode:2005GSLSP.252..279G. doi:10.1144/GSL.SP.2005.252.01.14.
3. Motani, Ryosuke (2005-05-31). "EVOLUTION OF FISH-SHAPED REPTILES (REPTILIA: ICHTHYOPTERYGIA) IN THEIR PHYSICAL ENVIRONMENTS AND CONSTRAINTS". Annual Review of Earth and Planetary Sciences. 33 (1): 395–420. doi:10.1146/annurev.earth.33.092203.122707. ISSN 0084-6597.
4. Fernández M. 2007. Redescription and phylogenetic position of Caypullisaurus (Ichthyosauria: Ophthalmosauridae). Journal of Paleontology 81 (2): 368-375.
5. Valentin Fischer; Michael W. Maisch; Darren Naish; Ralf Kosma; Jeff Liston; Ulrich Joger; Fritz J. Krüger; Judith Pardo Pérez; Jessica Tainsh; Robert M. Appleby (2012). "New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary". PLOS ONE. 7 (1): e29234. Bibcode:2012PLoSO...729234F. doi:10.1371/journal.pone.0029234. PMC 3250416. PMID 22235274.
6. Nikolay G. Zverkov & Megan L. Jacobs (2021) . "Revision of Nannopterygius (Ichthyosauria: Ophthalmosauridae): reappraisal of the 'inaccessible' holotype resolves a taxonomic tangle and reveals an obscure ophthalmosaurid lineage with a wide distribution". Zoological Journal of the Linnean Society. 191 (1): 228–275. doi:10.1093/zoolinnean/zlaa028.

v t e Ichthyosauromorpha
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Subclass: Diapsida Clade: Neodiapsida Sauropsida see Sauropsida Ichthyosauromorpha see below↓
Ichthyosauromorpha Ichthyosauromorpha Baisesaurus Hupehsuchia Eohupehsuchus Nanchangosaurus Hupehsuchidae Hupehsuchus Parahupehsuchinae Eretmorhipis Parahupehsuchus Ichthyosauriformes Omphalosauridae Cartorhynchus Omphalosaurus Sclerocormus Ichthyopterygia Isfjordosaurus? Chaohusaurus Thaisaurus Eoichthyosauria Grippidia Parvinatator Utatsusaurus Grippiidae Gulosaurus Grippia Ichthyosauria see below↓
Ichthyosauria Ichthyosauria Quasianosteosaurus Tholodus? Xinminosaurus? Cymbospondylidae Cymbospondylus Phantomosaurus? Thalattoarchon? Xinminosaurus? Hueneosauria Cymbospondylidae? Mixosauridae Barracudasauroides Contectopalatus Mixosaurus Phalarodon Merriamosauria Besanosaurus Guizhouichthyosaurus Pessopteryx? Wimanius? Shastasauridae Besanosaurus? Guanlingsaurus? Guizhouichthyosaurus? Himalayasaurus? Ichthyotitan? Shastasaurus Shonisaurus? Euichthyosauria Auroroborealia? Californosaurus Callawayia Guizhouichthyosaurus? Mikadocephalus? Shonisaurus? Toretocnemidae? Qianichthyosaurus? Toretocnemus Parvipelvia see below↓ Nomina dubia Barracudasaurus Pachygonosaurus Pessosaurus Sphaerodontes Svalbardosaurus Tibetosaurus
Parvipelvia Parvipelvia Hudsonelpidia Macgowania Neoichthyosauria Dearcmhara Hauffiopteryx Suevoleviathan Temnodontosauroidea? Temnodontosaurus Leptonectidae? Eurhinosaurus? Excalibosaurus? Leptonectes Wahlisaurus? Thunnosauria Ichthyosauridae Ichthyosaurus Malawania? Protoichthyosaurus? Baracromia Chacaicosaurus? Stenopterygiidae Magnipterygius Stenopterygius Ophthalmosauria Argovisaurus Delphinosaurus Macropterygius Ophthalmosauridae Arthropterygius? Gengasaurus? Jabalisaurus Janusaurus? Myobradypterygius Nannopterygius? Palvennia? Thalassodraco? Ophthalmosaurinae Acamptonectes Baptanodon Mollesaurus Ophthalmosaurus Platypterygiidae Acuetzpalin Aegirosaurus Arthropterygius? Athabascasaurus? Brachypterygius Catutosaurus Cryopterygius? Grendelius Janusaurus? Kazakhstanosaurus Keilhauia? Muiscasaurus Palvennia? Parrassaurus Pervushovisaurus Sumpalla Sveltonectes Undorosaurus Platypterygiinae Caypullisaurus Kyhytysuka Leninia Maiaspondylus Platypterygius Plutoniosaurus Simbirskiasaurus Sisteronia
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• Early Cretaceous ichthyosaurs
• Late Jurassic ichthyosaurs
• Berriasian life
• Tithonian life
• Late Jurassic reptiles of South America
• Early Cretaceous reptiles of South America
• Cretaceous Argentina
• Jurassic Argentina
• Fossils of Argentina
• Neuquén Basin
• Fossil taxa described in 1997
• Ophthalmosauridae
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