Browse history interactively← Previous editContent deleted Content addedVisualWikitextInline

Revision as of 15:05, 23 October 2007 view source86.128.124.233 (talk) →Inception of Darwin's evolutionary theory← Previous edit		Latest revision as of 20:17, 7 January 2025 view source Dave souza (talk | contribs)Autopatrolled, Administrators48,751 edits →Responses to publication: ce
Line 1:		Line 1:
			{{Short description|English naturalist and biologist (1809–1882)}}
	{{featured article}}
			{{Other people}}
	{{for|other people of the same surname, and places and things named after Charles Darwin|Darwin}}
			{{pp|small=yes}}
	{{Infobox_Scientist
			{{pp-move}}
	| name = Charles Robert Darwin
			{{Featured article}}
	| image = Charles Darwin aged 51.jpg
			{{Use British English|date=October 2024}}
	| caption = At the age of 51, Charles Darwin had just published </br> '']''.
			{{Use dmy dates|date=October 2024}}
	| birth_date = {{birth date|1809|2|12|df=y}}
			{{Infobox person
	| birth_place = ], ], ], ]
			| name = Charles Darwin
	| death_date = {{death date and age|1882|4|19|1809|2|12|df=yes}}
			| honorific_suffix = {{post-nominals|country=GBR|size=100%|JP|FRS|FRGS|FLS|FZS}}
	| death_place = ], ], ]
			| image = Charles Darwin seated crop.jpg
	| residence = ]
			| alt = Three quarter length studio photo showing Darwin's characteristic large forehead and bushy eyebrows with deep set eyes, pug nose and mouth set in a determined look. He is bald on top, with dark hair and long side whiskers but no beard or moustache. His jacket is dark, with very wide lapels, and his trousers are a light check pattern. His shirt has an upright wing collar, and his cravat is tucked into his waistcoat which is a light fine checked pattern.
	| nationality = ]
			| caption = Darwin, {{circa|1854}}, when he was preparing '']''{{sfn|Freeman|2007|p=76}}
	| field = ]
			| birth_name = Charles Robert Darwin
	| work_institutions = ]
			| birth_date = {{birth date|df=y|1809|2|12}}
	| alma_mater = ] <br /> ]
			| birth_place = ], ], England
	| doctoral_advisor = ]
			| death_date = {{death date and age|1882|df=y|4|19|1809|2|12}}
	| doctoral_students =
			| death_place = ], ], ], England<!--Letter no. 3368 footnote; "spelling of the village name was variable. 'Down' was the commonest form up to the 1870s, and 'Downe' thereafter", and it was in Kent until 1965, (now in ])-->
	| known_for = ]<br />]
			| resting_place = ]
	| prizes = ] (1853)<br />] (1859)<br />] (1864)
			| education = {{ubli|
	| religion = ], though ] family background, ] after 1851.
			]|
	| footnotes =
			] (], ])<ref>{{cite news|url=https://www.telegraph.co.uk/news/uknews/5032354/Charles-Darwins-personal-finances-revealed-in-new-find.html|archive-url=https://web.archive.org/web/20171019230458/http://www.telegraph.co.uk/news/uknews/5032354/Charles-Darwins-personal-finances-revealed-in-new-find.html|url-status=dead|title=Charles Darwin's personal finances revealed in new find|date=22 March 2009|archive-date=19 October 2017|work=The Telegraph
			}}</ref>}}
			| known_for = ]
			| spouse = {{marriage|]|1839}}
			| children = 10, including ], ], ], ], ] and ]
			| parents = {{ubl|]|]}}
			| family = ]
			| awards = {{ubl|] (1839)<ref name="catalogues.royalsociety.org-2015">{{cite web | title=Search Results: Record – Darwin; Charles Robert | website=The Royal Society Collections Catalogues | date=20 June 2015 | url=https://catalogues.royalsociety.org/CalmView/Record.aspx?src=CalmView.Persons&id=NA8196&pos=1 | access-date=2 December 2021 | archive-date=2 December 2021 | archive-url=https://web.archive.org/web/20211202200044/https://catalogues.royalsociety.org/CalmView/Record.aspx?src=CalmView.Persons&id=NA8196&pos=1 | url-status=live }}</ref>|] (1853)<ref name="Freeman 2007">{{harvnb|Freeman|2007|p=}}.</ref>|] (1859)<ref name="Freeman 2007" />|] (1864)<ref name="Freeman 2007" />|{{lang|fr|]}} (1867)<ref name="Freeman 2007" />|] (1879)<ref name="Freeman 2007" />}}
			| module = {{Infobox writer
			| embed = yes
			| notableworks = {{plainlist|
			* '']''
			* '']''
			* '']''}}
			| module = {{Infobox scientist
			| embed = yes
			| fields = {{ubli|]|]}}
			| work_institutions = ]
			| academic_advisors = {{ubl|]|]}}
			| author_abbrev_bot = '''Darwin'''
			| author_abbrev_zoo = '''Darwin'''
			| signature = Charles Darwin Signature.svg
			| signature_alt = "Charles Darwin", with the surname underlined by a downward curve that mimics the curve of the initial "C"
			}}
			}}
	}}		}}
	'''Charles Robert Darwin''' (] ] &ndash; ] ]) was an ] ].{{Ref_label|A|I|none}} After becoming eminent among scientists for his field work and inquiries into ], he proposed and provided ] evidence that all ] of life have ] over time from one or a few ]s through the process of ].<ref name=JvW>{{Harvnb|van Wyhe|2006}}.</ref> The ] became accepted by the ] and the general public in his lifetime, while his ] of natural selection came to be widely seen as the primary explanation of the process of evolution in the 1930s,<ref name=JvW>{{Harvnb|van Wyhe|2006}}.</ref> and now forms the basis of ]. In modified form, Darwin's scientific discovery remains the foundation of ], as it provides a unifying ] explanation for the ].<ref> ''darwin-online.org.uk''. Retrieved on ].<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Dobzhansky|1973}}</ref>
	<!--Please consider discussing changes on the talk page, as this opening is the result of a very long consensus-building process.-->		<!--Please consider discussing changes on the talk page, as this opening is the result of a very long consensus-building process.-->
			'''Charles Robert Darwin''' ({{IPAc-en|ˈ|d|ɑr|w|ɪ|n}}<ref> {{webarchive|url=https://web.archive.org/web/20140718234042/http://www.collinsdictionary.com/dictionary/english/darwin |date=18 July 2014 }} entry in '']''.</ref> {{Respell|DAR|win}}; 12 February 1809&nbsp;– 19 April 1882) was an English ], ], and ],<ref>{{harvnb|Desmond|Moore|Browne|2004}}.</ref> widely known for his contributions to ]. His proposition that all species of life have descended from a ] is now generally accepted and considered a fundamental scientific concept.<ref>{{cite book |last=Coyne |first=Jerry A. |author-link=Jerry Coyne |title=Why Evolution is True |publisher=Viking |year=2009 |pages= |isbn=978-0-670-02053-9 |url=https://archive.org/details/whyevolutionistr00coyn/page/8}}</ref> In a joint presentation with ], he introduced his scientific theory that this ] of ] resulted from a process he called ], in which the ] has a similar effect to the artificial selection involved in ].<ref name="Larson 2004">{{Harvnb|Larson|2004|pp=79–111}}.</ref> Darwin has been described as one of the most influential figures in ] and was honoured by ].<ref>{{cite news |url=https://www.newscientist.com/special/darwin-200 |title=Special feature: Darwin 200 |access-date=2 April 2011 |work=New Scientist |url-status=live |archive-url=https://web.archive.org/web/20110211051412/http://www.newscientist.com/special/darwin-200 |archive-date=11 February 2011}}</ref><ref name="Westminster Abbey-2016" />
	Darwin developed his interest in natural history while studying first ] at ], then ] at ].<ref name=whowas>{{Harvnb|Leff|2000}}.</ref> His ] on the ] established him as a ] whose observations and theories supported ]'s ] ideas, and publication of his ] made him famous as a popular author. Puzzled by the geographical distribution of wildlife and ]s he collected on the voyage, Darwin investigated the ] and conceived his theory of natural selection in 1838. Having seen others attacked as ] for such ideas, he confided only in his closest friends and continued his extensive research to meet anticipated objections.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 210, 263–274, 284–287}}.</ref> In 1858, ] sent him an essay describing a similar theory, causing the two to publish their theories early in a joint publication.<ref> ]. Retrieved on ].</ref>
			Darwin's early interest in nature led him to neglect his medical education at the ]; instead, he helped to investigate ]. His studies at the ]'s ] from 1828 to 1831 encouraged his passion for ].<ref name="Leff 2000">{{Harvnb|Leff|2000|loc=}}.</ref> However, it was his ] on {{HMS|Beagle}} from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported ]'s ]. Publication of his ] made Darwin famous as a popular author.<ref>{{Harvnb|Desmond|Moore|1991|pp=210, 284–285}}.</ref>
	His 1859 book '']'' established evolution by ] as the dominant scientific explanation of diversification in nature. He examined ] and ] in '']'', followed by '']''. His research on plants was published in a series of books, and in his final book, he examined ]s and their effect on soil.<ref>{{Harvnb|Freeman|1977}}</ref>		Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection.<ref>{{Harvnb|Desmond|Moore|1991|pp=263–274}}.</ref> Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority.<ref>{{harvnb|van Wyhe|2007|pp=184, 187}}</ref> He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting the immediate joint submission of ] to the ].<ref>{{Cite journal | last1=Beddall | first1=B. G. | title=Wallace, Darwin, and the Theory of Natural Selection | journal=Journal of the History of Biology | volume=1 | issue=2 | pages=261–323 | year=1968 | doi=10.1007/BF00351923 | s2cid=81107747 | df=dmy-all |issn=0022-5010}}</ref> Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification.<ref name="van Wyhe 2008" /> In 1871, he examined ] and ] in '']'', followed by '']'' (1872). His research on plants was published in a series of books, and in his final book, ] (1881), he examined ]s and their effect on soil.
			Darwin published his theory of evolution with compelling evidence in his 1859 book '']''.<ref>{{cite book |title=Why Evolution is True |last=Coyne |first=Jerry A. |author-link=Jerry Coyne |year=2009 |publisher=Oxford University Press |location=Oxford |isbn=978-0-19-923084-6 |page= |quote=In ''The Origin'', Darwin provided an alternative hypothesis for the development, diversification, and design of life. Much of that book presents evidence that not only supports evolution but at the same time refutes creationism. In Darwin's day, the evidence for his theories was compelling but not completely decisive. |url=https://archive.org/details/isbn_9780199230846/page/17 }}</ref><ref>{{cite book |title=Forerunners of Darwin |last=Glass |first=Bentley |author-link=Bentley Glass |year=1959 |publisher=Johns Hopkins University Press |location=Baltimore, MD |isbn=978-0-8018-0222-5 |page=iv |quote=Darwin's solution is a magnificent synthesis of evidence&nbsp;... a synthesis&nbsp;... compelling in honesty and comprehensiveness}}.</ref> By the 1870s, the scientific community and a majority of the educated public had accepted ]. However, many initially favoured ] that gave only a minor role to natural selection, and it was not until the emergence of the ] from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution.<ref name="van Wyhe 2008">{{Harvnb|van Wyhe|2008}}.</ref><ref name="Bowler 2003">{{harvnb|Bowler|2003|pp=178–179, 338, 347}}.</ref> Darwin's scientific discovery is the unifying theory of the ], explaining the ].
	In recognition of Darwin's pre-eminence, he was buried in ], close to ] and ].<ref>{{Harvnb|Browne|2002|p=497}}.</ref>
	==Biography==		==Biography==
	===Early life===
	{{details|Charles Darwin's education}}
			===Early life and education===
	]
			{{further|Charles Darwin's education|Darwin–Wedgwood family}}
			Darwin was born in ], Shropshire, on 12 February 1809, at his family's home, ].<ref>{{cite encyclopedia |last=Desmond |first=Adrian J. |title=Charles Darwin |encyclopedia=Encyclopædia Britannica |date=13 September 2002 |url=https://www.britannica.com/biography/Charles-Darwin |access-date=11 February 2018 |archive-date=6 February 2018 |archive-url=https://web.archive.org/web/20180206114419/https://www.britannica.com/biography/Charles-Darwin |url-status=live}}</ref><ref>{{cite web |url=http://darwin.baruch.cuny.edu/biography/shrewsbury/mount/ |title=The Mount House, Shrewsbury, England (Charles Darwin) |author=John H. Wahlert |date=11 June 2001 |work=Darwin and Darwinism |publisher=] |access-date=26 November 2008 |url-status=dead |archive-url=https://web.archive.org/web/20081206010149/http://darwin.baruch.cuny.edu/biography/shrewsbury/mount/ |archive-date=6 December 2008}}</ref> He was the fifth of six children of wealthy society doctor and financier ] and ] (née Wedgwood). His grandfathers ] and ] were both prominent ]. Erasmus Darwin had praised general concepts of evolution and ] in his '']'' (1794), a poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded.<ref>{{cite book |last=Smith |first=Homer W. |url=https://archive.org/details/manhisgods00smit |title=Man and His Gods |date=1952 |publisher=] |location=New York |pages= |author-link=Homer W. Smith |url-access=registration}}</ref>
			]. Part of ] showing him together with his sister Catherine.]]
	Charles Robert Darwin was born in ], ], ] on ] ] at his family home, ].<ref>, . Retrieved on ].</ref> He was the fifth of six children of wealthy society doctor and financier ], and ] (''née'' Wedgwood). He was the grandson of ] on his father's side, and of ] on his mother's side. Both families were largely ], though the Wedgwoods were adopting ]. Robert Darwin, himself quietly a ], made a nod toward convention by having baby Charles ] in the Anglican Church. Nonetheless, Charles and his siblings attended the Unitarian chapel with their mother, and in 1817, Charles joined the day school, run by its preacher. In July of that year, when Charles was eight years old, his mother died. From September 1818, he attended the nearby Anglican ] as a ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 12–15}}.</ref>
			Both families were largely ], though the Wedgwoods were adopting ]. Robert Darwin, a ], had baby Charles ] in November 1809 in the Anglican ], but Charles and his siblings attended the ] with their mother. The eight-year-old Charles already had a taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother ] in attending the nearby Anglican ] as a ].<ref name="Desmond-3">{{Harvnb|Desmond|Moore|1991|pp=12–15}};<br />{{harvnb|Darwin|1958|pp=}}.</ref>
	Darwin spent the summer of 1825 helping his father treat the poor of Shropshire as an apprentice doctor. In the autumn, he went to the ], Scotland, to study medicine, but he was revolted by the brutality of ] and neglected his medical studies. He learned ] from ], a freed black slave who told him exciting tales of the ]n ]. Later, in '']'', he used this experience as evidence that "Negroes and Europeans" were closely related despite superficial differences in appearance.<ref>{{Harvnb|Darwin|1871|loc=}}.</ref>		Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded ] with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies.{{sfn|Darwin|1958|pp=}} He learned ] in around 40 daily hour-long sessions from ], a freed black slave who had accompanied ] in the South American ].<ref name="Darwin 1958-6">{{harvnb|Darwin|1958|p=}};<br />{{harvnb|Desmond|Moore|2009|pp=}}.</ref>
	In Darwin's second year, he joined the Plinian Society, a student group interested in ].<ref>{{Harvnb|Browne|1995|p=72}}.</ref> He became a keen pupil of ], a proponent of ]'s ] by ], which Charles's grandfather Erasmus had also advocated. On the shores of the ], Darwin joined in Grant's investigations of the life cycle of ]. These studies found evidence for '']'', the ] theory that all animals have similar ] which differ only in complexity, thus showing ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p=33–40}}.</ref> In March 1827, Darwin made a presentation to the Plinian of his own discovery that the black spores often found in ] shells were the eggs of a skate ].<ref>{{Harvnb|Browne|1995|p=82}}.</ref> He also sat in on ]'s natural history course, learning about ] ], receiving training in ] plants, and assisting with work on the extensive collections of the ], one of the largest museums in Europe at the time.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p=42–43}}.</ref>		In Darwin's second year at the university, he joined the ], a student ] group featuring lively debates in which ] students with ] views challenged orthodox religious concepts of science.{{sfn|Desmond|Moore|1991|pp=31–34}} He assisted ]'s investigations of the anatomy and life cycle of ] in the ], and on 27 March 1827 presented at the Plinian his own discovery that black spores found in ] shells were the eggs of a skate ]. One day, Grant praised ]'s ]. Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals.<ref>{{Harvnb|Browne|1995|pp=72–88}}.</ref> Darwin was rather bored by ]'s natural-history course, which covered geology{{snd}}including the debate between ] and ]. He learned the ] of plants and assisted with work on the collections of the ], one of the largest museums in Europe at the time.<ref>{{Harvnb|Desmond|Moore|1991|pp=42–43}}.</ref>
			Darwin's neglect of medical studies annoyed his father, who sent him to ], in January 1828, to study for a Bachelor of Arts degree as the first step towards becoming an Anglican country ]. Darwin was unqualified for Cambridge's '']'' exams and was required instead to join the ordinary degree course.<ref>{{Harvnb|Browne|1995|pp=47–48, 89–91}};<br />{{harvnb|Desmond|Moore|2009|pp=47–48}}.</ref> He preferred ] and ] to studying.{{sfn|Darwin|1887|p=}}
	In 1827, his father, unhappy at his younger son's lack of progress, shrewdly enrolled him in a ] course at ] to qualify as a clergyman, expecting him to get a good income as an Anglican ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 47–48}}</ref> However, Darwin preferred ] and ] to studying.<ref>{{Harvnb|Darwin|1887|pp=, , , }}.</ref> Along with his cousin ], he became engrossed in the craze at the time for the competitive collecting of ]s.<ref>{{Harvnb|Darwin|1887|p=}}</ref> Fox introduced him to the Reverend ], professor of ], for expert advice on beetles. Darwin subsequently joined Henslow's natural history course and became his favourite pupil, known to the ] as "the man who walks with Henslow".<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 80–81}}</ref><ref>{{Harvnb|Darwin|1887|p=}}.</ref> When exams drew near, Darwin focused on his studies and received private instruction from Henslow. Darwin was particularly enthusiastic about the writings of ], including the ].<ref>{{Harvnb|Darwin|1887|p=}}</ref> It has been argued that Darwin's enthusiasm for Paley's religious adaptationism paradoxically played a role even later, when Darwin formulated his theory of natural selection.<ref>{{Harvnb|von Sydow|2005}}</ref>
	In his finals in January 1831, he performed well in ] and, having scraped through in ], ] and ], came tenth out of a pass list of 178.<ref>{{Harvnb|Browne|1995|p=97}}</ref>
			] portrait by ] of Darwin as a student, in the courtyard at ], where he had rooms.<ref name="BBC News-2009">{{cite web | title=Darwin statue unveiled at college | website=BBC News | date=12 February 2009 | url=http://news.bbc.co.uk/1/hi/england/cambridgeshire/7886278.stm | access-date=22 April 2022 | archive-date=16 February 2009 | archive-url=https://web.archive.org/web/20090216155859/http://news.bbc.co.uk/1/hi/england/cambridgeshire/7886278.stm | url-status=live }}</ref>]]
	Residential requirements kept Darwin at Cambridge until June. Following Henslow's example and advice, he was in no rush to take ]. Inspired by ]'s ''Personal Narrative'', he planned to visit the ] with some classmates after graduation to study natural history in the ]. To prepare himself, Darwin joined the geology course of the Reverend ] and, in the summer, went with him to assist in mapping strata in ].<ref>{{Harvnb|Browne|1995|pp= 133–141}}.</ref> After a fortnight with student friends at ], he returned home to find a letter from Henslow recommending Darwin as a suitable (if unfinished) naturalist for the unpaid position of gentleman's companion to ], the captain of ], which was to leave in four weeks on an expedition to chart the coastline of South America. His father objected to the planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, ], to agree to his son's participation.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 94–97}}.</ref>
			During the first few months of Darwin's enrolment at Christ's College, his second cousin ] was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to ] and influencing him to pursue ] collecting.<ref name="Smith-1952">{{cite book |last=Smith |first=Homer W. |url=https://archive.org/details/manhisgods00smit |title=Man and His Gods |date=1952 |publisher=] |location=New York |pages= |author-link=Homer W. Smith |url-access=registration}}</ref><ref name="Darwin 1887">{{harvnb|Darwin|1887|pp=}}.</ref> He did this zealously and had some of his finds published in ]' ''Illustrations of British entomology'' (1829–1932).<ref name="Darwin 1887" /><ref>{{Cite web |editor-last=van Wyhe |editor-first=John |title=Darwin's insects in Stephens' Illustrations of British entomology (1829–32) |url=http://darwin-online.org.uk/EditorialIntroductions/vanWyhe_Stephens.html |url-status=dead |access-date=3 July 2020 |website=Darwin Online |archive-date=1 September 2019 |archive-url=https://web.archive.org/web/20190901090213/http://darwin-online.org.uk/EditorialIntroductions/vanWyhe_Stephens.html}}</ref>
	===Journey of the ''Beagle''===
	{{details|Second voyage of HMS Beagle}}
			Through Fox, Darwin became a close friend and follower of botany professor ].<ref name="Smith-1952" /> He met other leading ]s who saw scientific work as religious ], becoming known to these ] as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of ]'s ''Evidences of Christianity'' (1795).<ref name="Desmond">{{Harvnb|Desmond|Moore|1991|pp=73–79, 763}};<br />{{Harvnb|Darwin|1958|pp=}}.</ref> In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for the ''ordinary'' degree.<ref>{{Harvnb|Browne|1995|p=97}}.</ref>
	]
			Darwin had to stay at Cambridge until June 1831. He studied Paley's '']'' (first published in 1802), which made an ], explaining ] as God acting through ].<ref name="von Sydow 2005">{{Harvnb|von Sydow|2005|pp=5–7}}.</ref> He read ]'s new book, ''Preliminary Discourse on the Study of Natural Philosophy'' (1831), which described the highest aim of ] as understanding such laws through ] based on observation, and ]'s ''Personal Narrative'' of scientific travels in 1799–1804.<ref>{{ cite book | last=Daum | first=Andreas W.|author-link=Andreas Daum | year=2024 | title=Alexander von Humboldt: A Concise Biography | location=Trans. Robert Savage. Princeton, N.J. | publisher=Princeton University Press |pages=137–138 | isbn=978-0-691-24736-6 }}</ref> Inspired with "a burning zeal" to contribute, Darwin planned to visit ] with some classmates after graduation to study natural history in the tropics. In preparation, he joined ]'s geology course, then on 4 August travelled with him to spend a fortnight mapping ] in Wales.<ref name="Darwin 1958-5">{{Harvnb|Darwin|1958|pp=}}.</ref><ref>{{Harvnb|Browne|1995|pp=128–129, 133–141}}.</ref>
	The ''Beagle'' survey took five years, two-thirds of which Darwin spent on land. He carefully noted a rich variety of geological features, ]s and living organisms, and methodically collected an enormous number of specimens, many of them new to science.<ref name=JvW/> At intervals during the voyage he sent specimens to Cambridge together with letters about his findings, and these established his reputation as a naturalist. His extensive detailed notes showed his gift for theorising and formed the basis for his later work. The journal he originally wrote for his family, published as '']'', summarises his findings and provides social, political and ] insights into the wide range of people he met, both native and colonial.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 189–192, 198}}.</ref>
			===Survey voyage on HMS ''Beagle''===
	While on board the ship, Darwin suffered badly from seasickness.<ref>{{Harvnb|Browne|1995|pp=177–178}}.</ref> In October 1833 he caught a fever in ], and in July 1834, while returning from the Andes down to ], he fell ill and spent a month in bed.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 142, 157}}.</ref>
			{{further|Second voyage of HMS Beagle}}
			], 1831–1836]]
	Before they set out, FitzRoy gave Darwin the first volume of ]'s ''Principles of Geology'', which explained landforms as the outcome of gradual processes over huge periods of time.{{Ref_label|B|II|none}} On their first stop ashore at ], Darwin found that a white band high in the ] cliffs consisted of baked ] fragments and shells. This matched Lyell's concept of land slowly rising or falling, giving Darwin a new insight into the geological history of the island which inspired him to think of writing a book on geology.<ref>{{Harvnb|Browne|1995|pp=183–190}}</ref> He went on to make many more discoveries, some of them particularly dramatic.<ref name=JvW/> He saw stepped plains of shingle and seashells in ] as ]es, and after experiencing an earthquake in ] saw ]-beds stranded above high tide showing that the land had just been raised. High in the ] he saw several fossil trees that had grown on a sand beach, with seashells nearby. He theorised that coral ]s form on sinking volcanic mountains, and confirmed this when the ''Beagle'' surveyed the ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 160–168, 182}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1887|p= }}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1958|loc=}}</ref>
			After leaving Sedgwick in Wales, Darwin spent a few days with student friends at ]. He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded ] place on {{HMS|Beagle}} with captain ], a position for a ] rather than "a mere collector". The ship was to leave in four weeks on an expedition to chart the coastline of South America.<ref name="Peter Lucas-2010">{{cite web |url=http://darwin-online.org.uk/EditorialIntroductions/Lucas_Lowe_Journal.html |title=The recovery of time past: Darwin at Barmouth on the eve of the Beagle |author=Peter Lucas |date=1 January 2010 |website=Darwin Online |access-date=5 December 2021 |archive-date=11 April 2010 |archive-url=https://web.archive.org/web/20100411090308/http://darwin-online.org.uk/EditorialIntroductions/Lucas_Lowe_Journal.html |url-status=live }}</ref><ref>{{cite web|url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-105.xml|title=Letter no. 105, Henslow, J. S. to Darwin, C. R., 24 Aug 1831|website=Darwin Correspondence Project|access-date=29 December 2021|archive-date=29 December 2021|archive-url=https://web.archive.org/web/20211229182043/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-105.xml|url-status=live}}</ref> Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, ], to agree to (and fund) his son's participation.<ref>{{Harvnb|Desmond|Moore|1991|pp=94–97}}</ref> Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution.<ref name="Browne 1995">{{Harvnb|Browne|1995|pp=204–210}}</ref>
	In South America, Darwin found and excavated rare fossils of gigantic extinct ]s in strata with modern seashells, indicating recent extinction and no change in climate or signs of catastrophe. Though he correctly identified one as a ] and fragments of armour reminded him of the local ], he assumed his finds were related to African or European species and it was a revelation to him after the voyage when ] showed that they were closely related to living creatures exclusively found in the Americas.<ref>{{Harvnb|Browne|1995|p= 224}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1835|p=}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 210}}</ref>
			After delays, the voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS ''Beagle'' ] coasts.<ref name="van Wyhe 2008" /><ref name="Keynes 2000">{{harvnb|Keynes|2000|pp=}}</ref> He kept careful notes of his observations and theoretical speculations. At intervals during the voyage, his specimens were sent to Cambridge together with letters including a copy of ] for his family.<ref>{{Harvnb|van Wyhe|2008b|pp=18–21}}</ref> He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal.<ref name="Randal Keynes-2006">{{cite web|url=http://darwin-online.org.uk/EditorialIntroductions/Chancellor_Keynes_Galapagos.html|title=Darwin's field notes on the Galapagos: 'A little world within itself'|author=Gordon Chancellor|author2=Randal Keynes|date=October 2006|publisher=]|access-date=16 September 2009|url-status=dead|archive-url=https://web.archive.org/web/20090901082402/http://darwin-online.org.uk/EditorialIntroductions/Chancellor_Keynes_Galapagos.html|archive-date=1 September 2009|author2-link=Randal Keynes}}</ref> Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with ] collected during a calm spell.<ref name="Keynes 2000" /><ref name="Keynes 2001-4">{{Harvnb|Keynes|2001|pp=}}</ref>
	] surveyed the coasts of ], Darwin began to theorise about the wonders of nature around him.]]
			] in Argentina, with fossils; caricature by ], the initial ship's artist]]
	Lyell's second volume, which argued against ] and explained species distribution by "centres of creation", was sent out to Darwin. He puzzled over all he saw, and his ideas went beyond Lyell.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 131, 159}}.</ref> In Argentina, he found that two types of ] had separate but overlapping territories. On the ], he collected ]s and noted that they were different depending on which island they came from. He also heard that local Spaniards could tell from their appearance on which island ]s originated, but thought the creatures had been imported by ]s.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 145, 170–172}}.</ref> In ], the ] ] and the ] seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work.<ref>{{Harvnb|Darwin|1839|p=}}.</ref>
			On their first stop ashore at ] in Cape Verde, Darwin found that a white band high in the ] cliffs included seashells. FitzRoy had given him the first volume of ]'s '']'', which set out ] concepts of land slowly rising or falling over immense periods,{{Ref label|B|II|none}} and Darwin saw things Lyell's way, theorising and thinking of writing a book on geology.<ref>{{Harvnb|Browne|1995|pp=183–190}}</ref> When they reached Brazil, Darwin was delighted by the ],<ref>{{harvnb|Keynes|2001|pp=}}</ref> but detested the sight of ], and disputed this issue with FitzRoy.<ref>{{harvnb|Darwin|1958|pp=}}</ref>
	In ] he and FitzRoy met ], who had recently written to Lyell about that "mystery of mysteries", the origin of species. When organising his notes on the return journey, Darwin wrote that if his growing suspicions about the mockingbirds and tortoises were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine".<ref>{{Harvnb|Keynes|2000}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Eldredge|2006}}</ref> He later wrote that such facts "seemed to me to throw some light on the origin of species".<ref>{{Harvnb|Darwin|1859|loc=}}</ref>
			The survey continued to the south in ]. They stopped at ], and in cliffs near ] Darwin made a major find of fossil bones of huge ] beside modern seashells, indicating recent ] with no signs of change in climate or catastrophe. He found bony plates like a giant version of the armour on local ]s. From a jaw and tooth he identified the gigantic '']'', then from ] description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest.<ref>{{Harvnb|Browne|1995|pp=223–225}}<br />{{Harvnb|Darwin|1835|p=}}<br />{{cite web|url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-213.xml|title=Letter no. 213, Henslow, J. S. to Darwin, C. R., 31 August 1833|website=Darwin Correspondence Project|access-date=29 December 2021|archive-date=29 December 2021|archive-url=https://web.archive.org/web/20211229182046/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-213.xml|url-status=live}}</ref><ref name="Keynes 2001">{{harvnb|Keynes|2001|pp=}}</ref> In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles.<ref name="Jaksic-2022">{{Cite journal |title=Historical account and current ecological knowledge of the southernmost lizard in the world, Liolaemus magellanicus (Squamata: Liolaemidae) |journal=] |last=Jaksic |first=Fabian M. |issue=7 |volume=95 |doi=10.1186/s40693-022-00112-y |year=2022|s2cid=252717680 |doi-access=free |bibcode=2022RvCHN..95....7J }}</ref>
	Three natives who had been taken from ] on the ''Beagle''<nowiki>'</nowiki>s previous voyage were taken back there to become missionaries.<!-- 4 were taken, one died in England, 3 returned: covered in Darwin's cite after next sentence<ref name=DarwinAsATraveller>{{Harvnb|Anonymous|1960}}</ref> --> They had become "civilised" in England over the previous two years, yet their relatives appeared to Darwin to be "miserable, degraded savages".<ref>{{Harvnb|Darwin|1845|pp= }}.</ref> A year on, the mission had been abandoned and only ] spoke with them to say he preferred his harsh previous way of life and did not want to return to England. Because of this experience, Darwin came to think that humans were not as far removed from animals as his friends then believed, and saw differences as relating to cultural advances towards civilisation rather than being racial. He detested the ] he saw elsewhere in South America, and was saddened by the effects of European settlement on ] in Australia and Maori in ].<ref>{{Harvnb|Browne|1995|p= 244–250}}</ref>
			On rides with ]s into the interior to explore geology and collect more fossils, Darwin gained social, political and ] insights into both native and colonial people at a time of revolution, and learnt that two types of ] had separate but overlapping territories.<ref>{{Harvnb|Desmond|Moore|1991|pp=189–192, 198}}</ref><ref>{{Harvnb|Eldredge|2006}}</ref> Further south, he saw stepped plains of shingle and seashells as ]es at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species.<ref>{{Harvnb|Desmond|Moore|1991|pp=131, 159}}<br />{{harvnb|Herbert|1991|pp=}}</ref><ref name="Darwin Online-2">{{cite web|url=http://darwin-online.org.uk/EditorialIntroductions/Chancellor_fieldNotebooks1.8.html|title=Darwin Online: 'Hurrah Chiloe': an introduction to the Port Desire Notebook|access-date=24 October 2008|url-status=dead|archive-url=https://web.archive.org/web/20081204201413/http://www.darwin-online.org.uk./EditorialIntroductions/Chancellor_fieldNotebooks1.8.html|archive-date=4 December 2008}}</ref>
	Captain FitzRoy was committed to writing the official ''Narrative'' of the ''Beagle'' voyages, and near the end of the voyage, he read Darwin's diary and asked him to rewrite this ''Journal'' to provide the third volume, on natural history.<ref>{{Harvnb|Browne|1995|p= 336}}</ref>
			Three ] on board, who had been seized during the ] then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at ] he met "miserable, degraded savages", as different as wild from domesticated animals.<ref>{{Harvnb|Darwin|1845|pp=}}</ref> He remained convinced that, despite this diversity, all humans were interrelated with ] and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there was no unbridgeable gap between humans and animals.<ref>{{Harvnb|Browne|1995|pp=243–244, 248–250, 382–383}}</ref> A year on, the mission had been abandoned. The Fuegian they had named ] lived like the other natives, had a wife, and had no wish to return to England.<ref>{{harvnb|Keynes|2001|pp=}}</ref>
			] surveyed the coasts of South America, Darwin theorised about geology and the extinction of giant mammals; watercolour by the ship's artist ], who replaced Augustus Earle, in ]]]
			Darwin experienced ] in 1835 and saw signs that the land had just been raised, including ]-beds stranded above high tide. High in the ] he saw seashells and several fossil trees that had grown on a sand beach. He theorised that as the land rose, ] sank, and ]s round them grew to form ]s.<ref>{{Harvnb|Desmond|Moore|1991|pp=160–168, 182}}<br />{{cite web | title=Letter no. 275 – Charles Darwin to Susan Elizabeth Darwin – 23 April 1835 | website=Darwin Correspondence Project | url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-275.xml | access-date=6 December 2021 | archive-date=6 December 2021 | archive-url=https://web.archive.org/web/20211206101229/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-275.xml | url-status=live }}</ref><ref name="Darwin 1958">{{Harvnb|Darwin|1958|loc=}}</ref>
			On the geologically new ], Darwin looked for evidence attaching wildlife to an older "centre of creation", and found ]s allied to those in Chile but differing from island to island. He heard that slight variations in the shape of ] shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food.<ref name="Keynes 2001-2">{{harvnb|Keynes|2001|pp=}}</ref><ref>{{harvnb|Sulloway|1982|p=19}}</ref> In Australia, the ] ] and the ] seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work.<ref name="Darwin Online">{{cite web|url=http://darwin-online.org.uk/EditorialIntroductions/Chancellor_fieldNotebooks1.3.html|title=Darwin Online: Coccatoos & Crows: An introduction to the Sydney Notebook|access-date=2 January 2009|url-status=live|archive-url=https://web.archive.org/web/20090114015611/http://darwin-online.org.uk/EditorialIntroductions/Chancellor_fieldNotebooks1.3.html|archive-date=14 January 2009}}</ref> He found the ] "good-humoured & pleasant", their numbers depleted by European settlement.<ref>{{harvnb|Keynes|2001|pp=.}}</ref>
			FitzRoy investigated how the atolls of the ] had formed, and the survey supported Darwin's theorising.<ref name="Darwin 1958" /> FitzRoy began writing the official ''Narrative'' of the ''Beagle'' voyages, and after reading Darwin's diary, he proposed incorporating it into the account.<ref name="Darwin Correspondence Project-5">{{cite web | title=Letter no. 301, Charles Darwin to Caroline Darwin, 29 April 1836, Port Lewis, Mauritius. | website=Darwin Correspondence Project | url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-301.xml | access-date=12 February 2022 | archive-date=15 February 2022 | archive-url=https://web.archive.org/web/20220215113535/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-301.xml | url-status=live }}</ref> Darwin's ''Journal'' was eventually rewritten as a separate third volume, on geology and natural history.<ref>{{Harvnb|Browne|1995|p=336}}</ref><ref name="Darwin 1839">{{Harvnb|Darwin|1839|p=}}</ref>
			In ], South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his ] as opening bold speculation on "that mystery of mysteries, the replacement of extinct species by others" as "a natural in contradistinction to a miraculous process".<ref name="van Wyhe 2007">{{harvnb|van Wyhe|2007|p=}}</ref>
			When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the ] were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine".<ref name="Keynes 2000-2">{{Harvnb|Keynes|2000|pp=}}<br />{{Harvnb|Eldredge|2006}}</ref> He later wrote that such facts "seemed to me to throw some light on the origin of species".<ref>{{Harvnb|Darwin|1859|loc=}}</ref>
			Without telling Darwin, ] had been read to scientific societies, printed as a pamphlet for private distribution among members of the ], and reported in magazines,{{sfn|Darwin|1835|p=1}} including ].<ref name="Darwin Correspondence Project-3">{{cite web | title=Letter no. 291, Caroline Darwin to Charles Darwin, 29 December , | website=Darwin Correspondence Project | url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-291.xml | access-date=19 January 2022 | archive-date=11 February 2022 | archive-url=https://web.archive.org/web/20220211205926/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-291.xml | url-status=live }}</ref> Darwin first heard of this at Cape Town,<ref name="Darwin Correspondence Project-4">{{cite web | title=Letter no. 302, Charles Darwin to Catherine Darwin, 3 June 1836, Cape of Good Hope | website=Darwin Correspondence Project | url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-302.xml | access-date=19 January 2022 | archive-date=27 January 2022 | archive-url=https://web.archive.org/web/20220127212812/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-302.xml | url-status=live }}</ref> and at ] read of Sedgwick's prediction that Darwin "will have a great name among the Naturalists of Europe".<ref name="Darwin Correspondence Project-2">{{cite web | title=Letter no. 288, Susan Darwin to Charles Darwin, 22 November 1835, Shrewsbury | website=Darwin Correspondence Project | url=https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-288.xml | access-date=19 January 2022 | archive-date=13 February 2022 | archive-url=https://web.archive.org/web/20220213100832/https://www.darwinproject.ac.uk/letter/?docId=letters/DCP-LETT-288.xml | url-status=live }}</ref><ref>{{harvnb|Darwin|1958|pp=.}}</ref>
	===Inception of Darwin's evolutionary theory===		===Inception of Darwin's evolutionary theory===
	{{details|Inception of Darwin's theory}}		{{further|Inception of Darwin's theory}}
			].]]
			On 2 October 1836, ''Beagle'' anchored at ], Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to ] to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be a self-funded ], and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage.<ref>{{Harvnb|Desmond|Moore|1991|pp=195–198}}</ref>
	]
			Charles Lyell eagerly met Darwin for the first time on 29 October and soon introduced him to the up-and-coming anatomist ], who had the facilities of the ] to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ]s as well as the ''Megatherium'' Darwin had identified, a near complete skeleton of the unknown '']'' and a ]-sized ]-like skull named '']'' resembling a giant ]. The armour fragments were actually from '']'', a huge armadillo-like creature, as Darwin had initially thought.<ref name="Keynes 2001" /><ref>{{Harvnb|Owen|1840|pp=, , }}<br />{{Harvnb|Eldredge|2006}}</ref> These extinct creatures were related to living species in South America.<ref>{{Harvnb|Desmond|Moore|1991|pp=201–205}}<br />{{Harvnb|Browne|1995|pp=349–350}}</ref>
	While Darwin was still on the voyage, ] fostered his former pupil's reputation by giving selected naturalists access to the fossil specimens and a pamphlet of Darwin's geological letters.<ref>{{Harvnb|Darwin|1835|loc= .}}</ref> When the ''Beagle'' returned on ] ], Darwin was a celebrity in scientific circles. After visiting his home in Shrewsbury and seeing relatives, Darwin hurried to ] to see Henslow, who advised on finding naturalists available to describe and catalogue the collections, and agreed to take on the botanical specimens. Darwin's father organised investments, enabling his son to be a self-funded ] scientist, and an excited Darwin went round the ] institutions being fêted and seeking experts to describe the collections. Zoologists had a huge backlog of work, and there was a danger of specimens just being left in storage.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 195–198}}.</ref>
			In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into the ''Narrative'' were resolved at the end of the month when FitzRoy accepted ] advice to make it a separate volume, and Darwin began work on his ''Journal and Remarks''.{{sfn|Browne|1995|pp=345–347}}{{sfn|Keynes|2001|pp=}}
	An eager ] met Darwin for the first time on ] and soon introduced him to the up-and-coming anatomist ], who had the facilities of the ] at his disposal to work on the fossil bones collected by Darwin. Owen's surprising results included gigantic ]s, a ]-like skull from the extinct ] '']'', and armour fragments from a huge extinct armadillo ('']''), as Darwin had initially surmised.<ref>{{Harvnb|Owen|1840|loc= }}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Eldredge|2006}}.</ref> The fossil creatures were unrelated to ]n animals, but closely related to living species in South America.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 201–205}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Browne|1995|p=349–350}}.</ref>
	In mid-December, Darwin moved to Cambridge to organise work on his collections and rewrite his ''Journal''.<ref>{{Harvnb|Browne|1995|p= 345–347}}.</ref> He wrote his first paper, showing that the South American landmass was slowly rising, and with Lyell's enthusiastic backing read it to the ] on ] ]. On the same day, he presented his mammal and bird specimens to the ]. The ornithologist ] soon revealed that the Galapagos birds that Darwin had thought a mixture of ], "]" and ]es, were, in fact, twelve ]. On ] ], Darwin was elected to the Council of the Geographical Society, and in his presidential address, Lyell presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his ] ideas.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 207–210}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Sulloway|1982|p=57}}</ref>		Darwin's first paper showed that the South American landmass was slowly rising. With Lyell's enthusiastic backing, he read it to the ] on 4 January 1837. On the same day, he presented his mammal and bird specimens to the ]. The ornithologist ] soon announced that the Galápagos birds that Darwin had thought a mixture of ], "]" and ]es, were, in fact, twelve ]. On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas.<ref>{{Harvnb|Desmond|Moore|1991|pp=207–210}}<br />{{Harvnb|Sulloway|1982|pp=20–23}}</ref>
	] from his ''First Notebook on Transmutation of Species'' (1837)]]		].]]
			Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as ],<ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-346.html|title=Darwin Correspondence Project&nbsp;– Letter 346&nbsp;– Darwin, C. R. to Darwin, C. S., 27 Feb 1837|access-date=19 December 2008|url-status=live|archive-url=https://web.archive.org/web/20090629192201/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-346.html|archive-date=29 June 2009}} proposes a move on Friday 3 March 1837,<br />Darwin's Journal ({{harvnb|Darwin|2006|pp=}}) backdated from August 1838 gives a date of 6 March 1837</ref> who described God as a programmer of laws. Darwin stayed with his ] brother Erasmus, part of this ] circle and a close friend of the writer ], who promoted the ] that underpinned the controversial Whig ] to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the ] implications of ], promoted by Grant and younger surgeons influenced by ]. Transmutation was anathema to Anglicans defending social order,<ref>{{Harvnb|Desmond|Moore|1991|pp=201, 212–221}}</ref> but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a ] of the origin of new species.<ref name="van Wyhe 2007" />
			Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought was a "]" was ]. Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands.<ref>{{Harvnb|Sulloway|1982|pp=9, 20–23}}</ref> The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards.<ref>{{Harvnb|Browne|1995|p=360}}<br />{{cite web|url=http://darwin-online.org.uk/content/frameset?itemID=F1643&viewtype=text&pageseq=1|title=Darwin, C. R. (Read 14 March 1837) Notes on Rhea americana and Rhea darwinii, ''Proceedings of the Zoological Society of London''|access-date=17 December 2008|url-status=live|archive-url=https://web.archive.org/web/20090210085710/http://darwin-online.org.uk/content/frameset?itemID=F1643&viewtype=text&pageseq=1|archive-date=10 February 2009}}</ref>
	On ] ], Darwin moved to London to be close to this work, and joined the social whirl around scientists and ]s such as ], who thought that God preordained life by ]s rather than ] miraculous creations. Darwin lived near his ] brother ], who was part of this ] circle and whose close friend the writer ] promoted the ideas of ] underlying the Whig "] reforms" aimed at discouraging the poor from breeding beyond available food supplies. ]'s question on the origin of species was widely discussed. Medical men including ] even joined ] in endorsing ], but to Darwin's scientist friends such ] heresy attacked the divine basis of the social order already under threat from recession and riots.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 196–201, 212–221}}.</ref>
			By mid-March 1837, barely six months after his return to England, Darwin was speculating in his ''Red Notebook'' on the possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal '']'', which resembled a giant ], a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations{{snd}}explaining the variations he had observed in ]s, mockingbirds, and rheas. He sketched branching descent, and then a ] branching of a single ], in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent ] progressing to higher forms.<ref>{{harvnb|Herbert|1980|pp=}}<br />{{Harvnb|van Wyhe|2008b|p=44}}<br />{{harvnb|Darwin|1837|pp=}}<br />{{Harvnb|Desmond|Moore|1991|pp=229–232}}</ref>
	Gould now revealed that the Galapagos ]s from different islands were separate species, not just varieties, and the "]s" were yet another species of finches. Darwin had not kept track of which islands the finch specimens were from, but found information from the notes of others on the ''Beagle'', including FitzRoy, who had more carefully recorded their own collections. The zoologist ] showed that the ]s were native to the islands. By mid-March, Darwin was convinced that creatures arriving in the islands had become altered in some way to form new species on the different islands, and investigated transmutation while noting his speculations in his "Red Notebook" which he had begun on the ''Beagle''. In mid-July, he began his secret "B" notebook on transmutation, and on page 36 wrote "I think" above his first sketch of an evolutionary tree.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 220–229}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Eldredge|2006}}.</ref>
	===Overwork, illness, and marriage ===		===Overwork, illness, and marriage===
			{{further|Health of Charles Darwin}}
	As well as launching into this intensive study of ], Darwin became mired in more work. While still rewriting his ''Journal'', he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained a Treasury grant of ]1,000 to sponsor this multi-volume '']''. He agreed to unrealistic dates for this and for a book on ''South American Geology'' supporting Lyell's ideas. Darwin finished writing his ''Journal'' around ] ] just as ] came to the throne, but then had its proofs to correct.<ref>{{Harvnb|Browne|1995|pp=367–369}}.</ref>
			While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his ''Journal'', he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained a Treasury grant of £1,000 to sponsor this multi-volume '']'', a sum equivalent to about £115,000 in 2021.<ref>{{cite web |title=£1,000 in 1832 → 2021 {{!}} UK Inflation Calculator |url=https://www.in2013dollars.com/uk/inflation/1832 |access-date=8 August 2021 |website=www.in2013dollars.com |archive-date=15 August 2021 |archive-url=https://web.archive.org/web/20210815234035/https://www.in2013dollars.com/uk/inflation/1832 |url-status=live }}</ref> He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher.<ref>{{Harvnb|Browne|1995|pp=367–369}}</ref> As the ] began, Darwin pressed on with writing his ''Journal'', and in August 1837 began correcting ].<ref name="Keynes 2001-3">{{harvnb|Keynes|2001|p=}}</ref>
	Darwin's health suffered from the pressure. On ] ], he had "palpitations of the heart". On doctor's advice that a month of recuperation was needed, he went to Shrewsbury then on to visit his Wedgwood relatives at ], but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin ], nine months older than Darwin, was nursing his invalid aunt. His uncle ] pointed out an area of ground where cinders had disappeared under ] and suggested that this might have been the work of ]s. This inspired a talk which Darwin gave to the Geological Society on ], the first demonstration of the role of earthworms in ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 233–234}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Arrhenius|1921|pp=255–257}}</ref>
			As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable ] of the heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at ], Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin ], nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under ] and suggested that this might have been the work of ]s, inspiring "a new & important theory" on their role in ], which Darwin presented at the Geological Society on 1 November 1837.<ref>{{Harvnb|Desmond|Moore|1991|pp=233–234}}<br />{{cite web |url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-404.html |title=Darwin Correspondence Project&nbsp;– Letter 404&nbsp;– Buckland, William to Geological Society of London, 9 Mar 1838 |access-date=23 December 2008 |url-status=live|archive-url=https://web.archive.org/web/20090629192234/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-404.html |archive-date=29 June 2009}}</ref> His ''Journal'' was printed and ready for publication by the end of February 1838, as was the first volume of the ''Narrative'', but FitzRoy was still working hard to finish his own volume.<ref name="Keynes 2001-3" />
	] pushed Darwin to take on the duties of Secretary of the Geological Society. After first declining this extra work, he accepted the post in March 1838.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 233–236}}.</ref> Despite the grind of writing and editing, remarkable progress was made on transmutation. While keeping his developing ideas secret, Darwin took every opportunity to question expert naturalists and, unconventionally, people with practical experience such as farmers and ].<ref name=JvW/><ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 241–244, 426}}.</ref> Over time his research drew on information from his relatives and children, the family butler, neighbours, colonists and former shipmates.<ref>{{Harvnb|Browne|1995|p=xii}}</ref> He included mankind in his speculations from the outset, and on seeing an ] in the zoo on ] ] noted its child-like behaviour.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 241–244}}.</ref>
			] pushed Darwin to take on the duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838.<ref>{{Harvnb|Desmond|Moore|1991|pp=233–236}}.</ref> Despite the grind of writing and editing the ''Beagle'' reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in ] such as farmers and ].<ref name="van Wyhe 2008" /><ref>{{Harvnb|Desmond|Moore|1991|pp=241–244, 426}}</ref> Over time, his research drew on information from his relatives and children, the family butler, neighbours, colonists and former shipmates.<ref>{{Harvnb|Browne|1995|p=xii}}</ref> He included mankind in his speculations from the outset, and on seeing an ] in the zoo on 28 March 1838 noted its childlike behaviour.<ref>{{Harvnb|Desmond|Moore|1991|pp=241–244}}</ref>
	The strain took its toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 252}}.</ref> For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe ]s, palpitations, trembling and other symptoms, particularly during times of stress, such as when attending meetings or dealing with controversy over his theory. The cause of ] was unknown during his lifetime, and attempts at treatment had little success. Recent attempts at diagnosis have suggested ] caught from insect bites in South America, ], or various psychological illnesses as possible causes, without any conclusive results.<ref>{{Harvnb|Gordon|Ref=CITEREFGordonThomas1999|1999}}.</ref>
			The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe ]s, palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of ] remained unknown, and attempts at treatment had only ephemeral success.<ref>{{Harvnb|Desmond|Moore|1991|pp=252, 476, 531}}<br />{{harvnb|Darwin|1958|p=}}</ref>
	On ] ], he took a break from the pressure of work and went "geologising" in Scotland. He visited ] in glorious weather to see the parallel "roads", horizontal ledges cut into the hillsides. He thought that these were ]es: they were later shown to have been shorelines of a ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 254}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Browne|1995|pp=377–378}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1887|p=}}</ref>
			On 23 June, he took a break and went "geologising" in Scotland. He visited ] in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a ].<ref>{{Harvnb|Desmond|Moore|1991|p=254}}<br />{{Harvnb|Browne|1995|pp=377–378}}<br />{{Harvnb|Darwin|1958|p=}}</ref>
	].]]
			].]]
	Fully recuperated, he returned to Shrewsbury in July. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about career and prospects on two scraps of paper, one with columns headed ''"Marry"'' and ''"Not Marry"''. Advantages included "constant companion and a friend in old age&nbsp;... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time."<ref>{{Harvnb|Darwin|1958|pp=}}</ref> Having decided in favour, he discussed it with his father, then went to visit Emma on ] ]. He did not get around to proposing, but against his father's advice he mentioned his ideas on transmutation.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 256–259}}.</ref>
			Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed ''"Marry"'' and ''"Not Marry"''. Advantages under "Marry" included "constant companion and a friend in old age&nbsp;... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time".<ref>{{Harvnb|Darwin|1958|pp=}}</ref> Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation.<ref>{{Harvnb|Desmond|Moore|1991|pp=256–259}}</ref>
			He married Emma on 29 January 1839 and they were the parents of ten children, seven of whom survived to adulthood.
			===Malthus and natural selection===
	Continuing his research in London, Darwin's wide reading now included "for amusement" the 6th edition of ] '']'' which calculates from the birth rate that human population could double every 25 years, but in practice growth is kept in check by death, disease, wars and famine.<ref name=JvW/><ref>{{Harvnb|Malthus|1826}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 264–265}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Huxley|1897|pp=}}</ref> Darwin was well prepared to see at once that this also applied to ] "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. This would result in the formation of new species.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 264–265}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Browne|1995|p= 385–388}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1842|p=}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1887|loc=}}</ref> On ] ] he noted this insight, describing it as a kind of wedging, forcing adapted structures into gaps in the economy of nature as weaker structures were thrust out.<ref name=JvW/> He now had a theory by which to work, and over the following months compared farmers picking the best breeding stock to a Malthusian Nature selecting from variants thrown up by "chance" so that "every part of &#91;every&#93; newly acquired structure is fully practised and perfected", and thought this analogy "the most beautiful part of my theory".<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 273–274}}.</ref>
			Continuing his research in London, Darwin's wide reading now included the sixth edition of ]'s '']''. On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a ] so that population soon exceeds food supply in what is known as a ]. Darwin was well-prepared to compare this to ]'s "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that the "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones."<ref name="van Wyhe 2008" /><ref name="Darwin transmutation notebook D pp">{{cite web | title=Darwin transmutation notebook D pp. 134e–135e | url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR123.-&pageseq=112 | access-date=4 June 2012 | url-status=live | archive-url=https://web.archive.org/web/20120718105154/http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR123.-&pageseq=112 | archive-date=18 July 2012 | df=dmy-all }}</ref> This would result in the formation of new species.<ref name="van Wyhe 2008" /><ref>{{Harvnb|Desmond|Moore|1991|pp=264–265}}<br />{{Harvnb|Browne|1995|pp=385–388}}<br />{{Harvnb|Darwin|1842|p=}}</ref> As he later wrote in his '']'':
	On ], he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness, but her upbringing as a very devout ] led her to express fears that his lapses of faith could endanger her hopes to meet in the afterlife.<ref>{{Harvnb|Browne|1995|p= 391–398}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 269–271}}.</ref> While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in ], then moved his "museum" in over Christmas. The marriage was arranged for ] ], but the Wedgwoods set the date back. On the 24th, Darwin was honoured by being elected as ] ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 272–279}}.</ref>
			{{Blockquote|In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate the struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work...<ref name="Darwin 1958-2">{{harvnb|Darwin|1958|p=}}</ref>}}
	On ] ], Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 279}}.</ref>
			By mid-December, Darwin saw a similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected",<ref>{{cite web |url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR124.-&pageseq=63 |title=Darwin transmutation notebook E p. 75 |access-date=17 March 2009 |url-status=live |archive-url=https://web.archive.org/web/20090628082830/http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR124.-&pageseq=63 |archive-date=28 June 2009 }}</ref> thinking this comparison "a beautiful part of my theory".<ref>{{cite web|url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR124.-&pageseq=61|title=Darwin transmutation notebook E p. 71|access-date=17 March 2009|url-status=live|archive-url=https://web.archive.org/web/20090628080656/http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=CUL-DAR124.-&pageseq=61|archive-date=28 June 2009}}</ref> He later called his theory ], an analogy with what he termed the "artificial selection" of selective breeding.<ref name="van Wyhe 2008" />
	===Preparing the theory of natural selection for publication===
	{{details|Development of Darwin's theory}}
			On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in the afterlife.<ref name="Darwin Correspondence Project">{{cite web|url=http://www.darwinproject.ac.uk/content/view/130/125/|title=Darwin Correspondence Project&nbsp;– Belief: historical essay|access-date=25 November 2008 |archive-url=https://web.archive.org/web/20090225124103/http://www.darwinproject.ac.uk/content/view/130/125/ |archive-date=25 February 2009 }}</ref> While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in ], then moved his "museum" in over Christmas. On 24 January 1839, Darwin was ] (FRS).<ref name="catalogues.royalsociety.org-2015">{{cite web | title=Search Results: Record – Darwin; Charles Robert | website=The Royal Society Collections Catalogues | date=20 June 2015 | url=https://catalogues.royalsociety.org/CalmView/Record.aspx?src=CalmView.Persons&id=NA8196&pos=1 | access-date=2 December 2021 | archive-date=2 December 2021 | archive-url=https://web.archive.org/web/20211202200044/https://catalogues.royalsociety.org/CalmView/Record.aspx?src=CalmView.Persons&id=NA8196&pos=1 | url-status=live }}</ref><ref>{{Harvnb|Desmond|Moore|1991|pp=272–279}}</ref>
	Darwin had found the basis of his theory of ], but was aware of how much work remained to make it credible to his fiercely critical scientific colleagues. As Secretary of the Geological Society at its meeting on ] ], he saw ] and ] display their hatred of evolution when destroying the reputation of his old ] teacher ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 274–276}}.</ref> Work on his ''Beagle'' findings continued, and as well as consulting ] he carried out extensive experiments with plants, trying to find evidence answering all the arguments he anticipated when his theory was made public.<ref>{{Harvnb|Darwin|1859|loc=ch. 1}}.</ref> When FitzRoy's ''Narrative'' was published in May 1839, Darwin's ''Journal and Remarks'' ('']'') as the third volume was such a success that later that year it was published on its own.<ref>{{Harvnb|Darwin|1887|loc=}}</ref>
			On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home.<ref>{{Harvnb|Desmond|Moore|1991|p=279}}</ref>
	Early in 1842, Darwin sent a letter about his ideas to ], who was dismayed that his ally now denied "seeing a beginning to each crop of species". In May, Darwin's book on ]s was published after more than three years of work, and he then wrote a "pencil sketch" of his theory.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 292}}.</ref> To escape the pressures of London, the family moved to rural ] in November.<ref>{{Harvnb|Darwin|1887|p=}}.</ref> On ] ] Darwin wrote to his botanist friend ] about his theory, saying it was like confessing "a murder", but to his relief Hooker thought that "there might have been a gradual change of species" and expressed interest in Darwin's explanation. By July, Darwin had expanded his "sketch" into a 230-page "Essay".<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 313–317}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1887|loc=}}</ref> His fears that his ideas would be dismissed as ] ] were reawakened by controversy over the anonymous publication in October of '']'', which was severely attacked by establishment scientists. However, the book was a best-seller and widened middle-class interest in transmutation, paving the way for Darwin as well as reminding him of the need to answer all difficulties before making his theory public. Darwin completed his third geological book in 1846, and embarked on a huge study of ]s with the assistance of Hooker. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 320–323, 339–348}}.</ref>
			===Geology books, barnacles, evolutionary research===
	In an attempt to improve his chronic ill health, Darwin went to a spa in ] in 1849. To his surprise, he found that two months of water treatment helped.<ref>{{Harvnb|Darwin|1887|p=}}</ref> Then his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. After a long series of crises, she died and Darwin ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 383–387}}.</ref>
			{{further|Development of Darwin's theory}}
			].]]
	Darwin's eight years of work on barnacles (''Cirripedia'') found "]" that supported his theory by showing that slightly changed body parts could serve different functions to meet new conditions.<ref>{{Harvnb|Darwin|1887|pp=,}}.</ref> In 1853 it earned him the ]'s Royal Medal, and it made his reputation as a ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 383–387}}.</ref> In 1854 he resumed work on his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in the economy of nature".<ref>{{Harvnb|Darwin|1887|pp=, }}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 419–420}}.</ref>
			Darwin now had the framework of his theory of natural selection "by which to work",<ref name="Darwin 1958-2" /> as his "prime hobby".<ref name="Darwin Correspondence Project-6">{{cite news |url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-419.html |title=Darwin Correspondence Project&nbsp;– Letter 419&nbsp;– Darwin, C. R. to Fox, W. D., (15 June 1838) |newspaper=Darwin Correspondence Project |access-date=8 February 2008 |url-status=live |archive-url=https://web.archive.org/web/20070904124133/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-419.html |archive-date=4 September 2007}}</ref> His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory.<ref name="van Wyhe 2008" /> For fifteen years this work was in the background to his main occupation of writing on geology and publishing expert reports on the ''Beagle'' collections, in particular, the ]s.<ref name="van Wyhe 2007-2">{{Harvnb|van Wyhe|2007|pp=}}</ref>
			The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed ]. His confusion over the relationship of this species (''Cryptophialus minutus'') to other barnacles caused him to fixate on the systematics of the taxa. He wrote his first examination of the species in 1846 but did not formally describe it until 1854.<ref name="buch">{{cite journal |last1=Buchanan |first1=Roderick D. |title=Darwin's "Mr. Arthrobalanus": Sexual Differentiation, Evolutionary Destiny and the Expert Eye of the Beholder |journal=Journal of the History of Biology |date=May 2017 |volume=50 |issue=2 |pages=315–355 |doi=10.1007/s10739-016-9444-9 |pmid=27098777 |url=https://pubmed.ncbi.nlm.nih.gov/27098777/ |access-date=3 September 2024 |issn=1573-0387}}</ref>
			FitzRoy's long-delayed ''Narrative'' was published in May 1839. Darwin's ''Journal and Remarks'' got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing a beginning to each crop of species".<ref name="Darwin 1839" /><ref>{{Harvnb|Desmond|Moore|1991|pp=284–285, 292}}</ref>
			Darwin's book '']'' on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection.<ref>{{Harvnb|Desmond|Moore|1991|pp=292–293}}<br />{{Harvnb|Darwin|1842|pp=}}</ref> To escape the pressures of London, the family moved to rural ] in Kent in September.<ref>{{Harvnb|Darwin|1958|p=}}</ref> On 11 January 1844, Darwin mentioned his theorising to the botanist ], writing with melodramatic humour "it is like confessing a murder".<ref>{{harvnb|van Wyhe|2007|pp=}}</ref><ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-729.html#back-mark-729.f6|title=Darwin Correspondence Project&nbsp;– Letter 729&nbsp;– Darwin, C. R. to Hooker, J. D., (11 January 1844)|access-date=8 February 2008|url-status=live|archive-url=https://web.archive.org/web/20080307235150/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-729.html#back-mark-729.f6|archive-date=7 March 2008}}</ref> Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject."<ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-734.html|title=Darwin Correspondence Project&nbsp;– Letter 734&nbsp;– Hooker, J. D. to Darwin, C. R., 29 January 1844|access-date=8 February 2008|url-status=live|archive-url=https://web.archive.org/web/20090226141303/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-734.html|archive-date=26 February 2009}}</ref>
			] in Kent was his usual "thinking path"<ref name="Darwin Online-3">{{cite web | title=Charles Darwin: a life in pictures, The Sand Walk near Down House, Darwin's thinking path | website=Darwin Online | url=http://darwin-online.org.uk/life19.html | access-date=1 October 2022 | archive-date=1 October 2022 | archive-url=https://web.archive.org/web/20221001173155/http://darwin-online.org.uk/life19.html | url-status=live }}</ref>]]
			By July, Darwin had expanded his "sketch" into a 230-page "Essay", to be expanded with his research results if he died prematurely.<ref>{{Harvnb|van Wyhe|2007|p=188}}</ref> In November, the anonymously published sensational best-seller '']'' brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists.<ref>{{harvnb|Browne|1995|pp=461–465}}</ref><ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-814.html#back-mark-814.f5|title=Darwin Correspondence Project&nbsp;– Letter 814&nbsp;– Darwin, C. R. to Hooker, J. D., (7 Jan 1845)|access-date=24 November 2008|url-status=live|archive-url=https://web.archive.org/web/20081205084645/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-814.html#back-mark-814.f5|archive-date=5 December 2008}}</ref>
			Darwin completed his third geological book in 1846. He now renewed a fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures.<ref>{{Harvnb|van Wyhe|2007|pp=190–191}}</ref> In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of ].<ref>{{Harvnb|Desmond|Moore|1991|pp=320–323, 339–348}}</ref>
			In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. ]'s ] spa and was surprised to find some benefit from ].<ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-1236.html|title=Darwin Correspondence Project&nbsp;– Letter 1236&nbsp;– Darwin, C. R. to Hooker, J. D., 28 Mar 1849|access-date=24 November 2008|url-status=live|archive-url=https://web.archive.org/web/20081207005457/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-1236.html|archive-date=7 December 2008}}</ref> Then, in 1851, his treasured daughter ] fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises.<ref>{{harvnb|Browne|1995|pp=498–501}}</ref>
			In eight years of work on barnacles, Darwin's theory helped him to find "]" showing that slightly changed body parts served different functions to meet new conditions, and in some ] he found minute males ] on ]s, showing an ] in evolution of ].<ref name="Darwin 1958-3">{{Harvnb|Darwin|1958|pp=}}</ref> In 1853, it earned him the ]'s Royal Medal, and it made his reputation as a ].<ref>{{Harvnb|Desmond|Moore|1991|pp=383–387}}</ref> Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before."<ref>{{Cite journal |last=Bromham |first=Lindell |date=1 October 2020 |title=Comparability in evolutionary biology: The case of Darwin's barnacles |url=https://www.degruyter.com/document/doi/10.1515/lingty-2020-2056/html?lang=en |journal=Linguistic Typology |language=en |volume=24 |issue=3 |pages=427–463 |doi=10.1515/lingty-2020-2056 |issn=1613-415X |s2cid=222319487 |hdl-access=free |hdl=1885/274303}}</ref>{{sfn|van Wyhe|2007}} In 1854, he became a Fellow of the ], gaining postal access to its library.<ref>{{harvnb|Freeman|2007|pp=107, 109}}</ref> He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in the economy of nature".<ref>{{Harvnb|Desmond|Moore|1991|pp=419–420}}</ref>
	===Publication of the theory of natural selection===		===Publication of the theory of natural selection===
	{{details|Publication of Darwin's theory}}		{{further|Publication of Darwin's theory}}
	].]]
			]. He wrote to ] about this portrait, "if I really have as bad an expression, as my photograph gives me, how I can have one single friend is surprising."<ref> {{webarchive|url=https://web.archive.org/web/20120107045842/http://darwin-online.org.uk/EditorialIntroductions/vanWyhe_MaullandPolyblankPhoto.html |date=7 January 2012 }}, John van Wyhe, December 2006</ref>]]
	By the start of 1856, Darwin was investigating whether eggs and ]s could survive travel across seawater to spread species across oceans. ] increasingly doubted the traditional view that species were fixed, but their young friend ] was firmly against evolution. ] was intrigued by Darwin's speculations without realising their extent. When he read a paper by ] on the ''Introduction'' of species, he saw similarities with Darwin's thoughts and urged him to publish to establish precedence. Though Darwin saw no threat, he began work on a short paper. Finding answers to difficult questions held him up repeatedly, and he expanded his plans to a "big book on species" titled ''Natural Selection''. He continued his researches, ] and specimens from naturalists worldwide including Wallace who was working in ]. In December 1857, Darwin received a letter from Wallace asking if the book would examine human origins. He responded that he would avoid that subject, "so surrounded with prejudices", while encouraging Wallace's theorising and adding that "I go much further than you."<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 412–441, 462–463}}.</ref>
			By the start of 1856, Darwin was investigating whether eggs and ]s could survive travel across seawater to spread species across oceans. Hooker increasingly doubted the traditional view that species were fixed, but their young friend ] was still firmly against the transmutation of species. Lyell was intrigued by Darwin's speculations without realising their extent. When he read a paper by ], "On the Law which has Regulated the Introduction of New Species", he saw similarities with Darwin's thoughts and urged him to publish to establish precedence.<ref name="Desmond-2">{{Harvnb|Desmond|Moore|1991|pp=412–441, 457–458, 462–463}}<br />{{harvnb | Desmond |Moore | 2009 | pp=283–284, 290–292, 295}}</ref>
	Darwin's book was half way when, on ] ], he received a paper from Wallace describing natural selection. Though shocked that he had been "forestalled", Darwin sent it on to Lyell, as requested, and, though Wallace had not asked for publication, offered to send it to any journal that Wallace chose. His family was in crisis with children in the village dying of ], and he put matters in the hands of Lyell and Hooker. They agreed on a joint presentation at the ] on ] of '']''; however, Darwin's baby son died of the scarlet fever and he was too distraught to attend.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 466–470}}.</ref>
			Though Darwin saw no threat, on 14 May 1856 he began writing a short paper. Finding answers to difficult questions held him up repeatedly, and he expanded his plans to a "big book on species" titled '']'', which was to include his "note on Man". He continued his research, ] and specimens from naturalists worldwide, including Wallace who was working in ].<ref name="Desmond-2" />
	There was little immediate attention to this announcement of the theory; the president of the Linnean remarked in May 1859 that the year had not been marked by any revolutionary discoveries.<ref>{{Harvnb|Browne|2002|p=40–42}}</ref> Later, Darwin could only recall one review; Professor Haughton of Dublin claimed that "all that was new in them was false, and what was true was old."<ref>{{Harvnb|Darwin|1958|p=}}.</ref> Darwin struggled for thirteen months to produce an abstract of his "big book", suffering from ill health but getting constant encouragement from his scientific friends. Lyell arranged to have it published by ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 374–474}}.</ref>
			In mid-1857, he added a section heading, "Theory applied to Races of Man", but did not add text on this topic. On 5 September 1857, Darwin sent the American botanist ] a detailed outline of his ideas, including an abstract of ''Natural Selection'', which omitted ] and ]. In December, Darwin received a letter from Wallace asking if the book would examine human origins. He responded that he would avoid that subject, "so surrounded with prejudices", while encouraging Wallace's theorising and adding that "I go much further than you."<ref name="Desmond-2" />
	''On the Origin of Species by Means of Natural Selection, or The Preservation of Favoured Races in the Struggle for Life'' (usually abbreviated to '']'') proved unexpectedly popular, with the entire stock of 1,250 copies oversubscribed when it went on sale to booksellers on ] ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 477}}.</ref> In the book, Darwin set out "one long argument" of detailed observations, inferences and consideration of anticipated objections.<ref>{{Harvnb|Darwin|1859|loc= }}</ref> His only allusion to human evolution was the understatement that "light will be thrown on the origin of man and his history".<ref>{{Harvnb|Darwin|1859|loc= }}</ref> He avoided the then controversial term "]", but at the end of the book concluded that "endless forms most beautiful and most wonderful have been, and are being, evolved."<ref>{{Harvnb|Darwin|1859|loc= }}</ref> His theory is simply stated in the introduction:
			Darwin's book was only partly written when, on 18 June 1858, he received a paper from Wallace describing natural selection. Shocked that he had been "forestalled", Darwin sent it on that day to Lyell, as requested by Wallace,<ref>Ball, P. (2011). Shipping timetables debunk Darwin plagiarism accusations: Evidence challenges claims that Charles Darwin stole ideas from Alfred Russel Wallace. Nature. {{webarchive|url=https://web.archive.org/web/20120222191430/http://www.nature.com/news/shipping-timetables-debunk-darwin-plagiarism-accusations-1.9613 |date=22 February 2012 }}</ref><ref>{{cite journal |doi=10.1111/j.1095-8312.2011.01808.x|title=A new theory to explain the receipt of Wallace's Ternate Essay by Darwin in 1858|journal=Biological Journal of the Linnean Society|volume=105|pages=249–252|year=2012|last1=van Wyhe|first1=John|last2=Rookmaaker|first2=Kees|doi-access=free}}</ref> and although Wallace had not asked for publication, Darwin suggested he would send it to any journal that Wallace chose. His family was in crisis, with children in the village dying of ], and he put matters in the hands of his friends. After some discussion, with no reliable way of involving Wallace, Lyell and Hooker decided on a joint presentation at the Linnean Society on 1 July of '']''. On the evening of 28 June, Darwin's baby son died of scarlet fever after almost a week of severe illness, and he was too distraught to attend.<ref>{{Harvnb|Desmond|Moore|1991|pp=466–470}}</ref>
	<blockquote>
	As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be ''naturally selected''. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.<ref>{{Harvnb|Darwin|1859|loc= }}</ref>
	</blockquote>
			There was little immediate attention to this announcement of the theory; the president of the Linnean Society remarked in May 1859 that the year had not been marked by any revolutionary discoveries.<ref>{{Harvnb|Browne|2002|pp=40–42, 48–49}}</ref> Only one review rankled enough for Darwin to recall it later; Professor ] of Dublin claimed that "all that was new in them was false, and what was true was old".<ref>{{Harvnb|Darwin|1958|p=}}</ref> Darwin struggled for thirteen months to produce an abstract of his "big book", suffering from ill health but getting constant encouragement from his scientific friends. Lyell arranged to have it published by ].<ref>{{Harvnb|Desmond|Moore|1991|pp=374–474}}</ref>
	]
			'']'' proved unexpectedly popular, with the entire stock of 1,250 copies oversubscribed when it went on sale to booksellers on 22 November 1859.<ref>{{Harvnb|Desmond|Moore|1991|p=477}}</ref> In the book, Darwin set out "one long argument" of detailed observations, inferences and consideration of anticipated objections.<ref>{{Harvnb|Darwin|1859|loc=}}</ref> In making the case for common descent, he included evidence of homologies between humans and other mammals.{{sfn|van Wyhe|2008}}{{Ref label|C|III|none}} Having outlined sexual selection, he hinted that it could explain differences between ].<ref name="Darwin 1859-2">{{Harvnb|Darwin|1859|p=}}<br />{{harvnb | Darwin |Costa | 2009 | p=199}}<br />{{harvnb | Desmond |Moore | 2009 | p=310}}</ref>{{Ref label|D|IV|1}} He avoided explicit discussion of human origins, but implied the significance of his work with the sentence; "Light will be thrown on the origin of man and his history."<ref name="Darwin 1859">{{harvnb|Darwin|1859|p=}}<br />{{harvnb | Darwin |Costa | 2009 | pp=199, 488}}<br />{{harvnb|van Wyhe|2008}}</ref>{{Ref label|D|IV|2}} His theory is simply stated in the introduction:
	===Reaction to the publication===
			{{blockquote|As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be ''naturally selected''. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.<ref>{{Harvnb|Darwin|1859|loc=}}</ref>}}
	{{details|Reaction to Darwin's theory}}
			At the end of the book, he concluded that:
	There was wide public interest in Charles Darwin's book and a controversy which he monitored closely, keeping press cuttings of ]s, ], ]s, ] and ]s.<ref>{{Harvnb|Browne|2002|p=103–104, 379}}</ref> Critical reviewers were quick to pick out the unstated implications of "men from ]s", while amongst favourable responses Huxley's reviews included swipes at ], leader of the scientific establishment Huxley was trying to overthrow. Owen's verdict was unknown until his April review condemned the book.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 477–491}}.</ref>
			{{blockquote|There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.<ref>{{Harvnb|Darwin|1859|loc=}}</ref>}}
			The last word was the only variant of "evolved" in the first five editions of the book. "]" at that time was associated with other concepts, most commonly with ]. Darwin first used the word ] in '']'' in 1871, before adding it in 1872 to the 6th edition of ''The Origin of Species''.<ref>{{harvnb|Browne|2002|p=59}}, {{harvnb|Freeman|1977|pp=}}</ref>
	The ] scientific establishment, including Darwin's old Cambridge tutors ] and ], reacted against the book, though it was well received by a younger generation of professional naturalists. In 1860, the publication of '']'' by seven liberal Anglican theologians diverted ] attention away from Darwin. An explanation of ] and other ], it included the argument that ]s broke God's laws, so belief in them was ]&mdash;and praise for "Mr Darwin's masterly volume &#91;supporting&#93; the grand principle of the self-evolving powers of nature".<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 487–488, 500}}.</ref>
			===Responses to publication===
	The most famous confrontation took place at a meeting of the ] in ]. Professor ] delivered a long lecture about Darwin and social progress, then ], the ], argued against Darwin. In the ensuing debate ] argued strongly for Darwin and ] established himself as "Darwin's bulldog" &ndash; the fiercest defender of evolutionary theory on the Victorian stage. Both sides came away feeling victorious, but Huxley went on to make much of his claim that on being asked by Wilberforce whether he was descended from monkeys on his grandfather's side or his grandmother's side, Huxley muttered: "The Lord has delivered him into my hands" and replied that he "would rather be descended from an ape than from a cultivated man who used his gifts of culture and eloquence in the service of prejudice and falsehood".<ref>{{Harvnb|Lucas|1979}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 493–499}}.</ref>
			{{further|Reactions to On the Origin of Species}}
			].<ref name="Browne 2002-2" />]]
			]'' was typical of many showing Darwin with an ] body, identifying him in popular culture as the leading author of evolutionary theory.<ref name="Browne 2002-2" />]]
			The book aroused international interest, with less controversy than had greeted the popular and less scientific ''Vestiges of the Natural History of Creation''.<ref>{{harvnb|van Wyhe|2008b|p=48}}</ref> Though Darwin's illness kept him away from the public debates, he eagerly scrutinised the scientific response, commenting on press cuttings, reviews, articles, satires and caricatures, and corresponded on it with colleagues worldwide.<ref>{{Harvnb|Browne|2002|pp=103–104, 379}}</ref> The book did not explicitly discuss human origins,<ref name="Darwin 1859" />{{Ref label|D|IV|3}} but included a number of hints about the animal ancestry of humans from which the inference could be made.<ref>{{harvnb|Radick|2013|pp=174–175}}<br />{{harvnb|Huxley|Kettlewell|1965|p=88}}</ref>
	Darwin's illness kept him away from the public debates, though he read eagerly about them and mustered support through ]. ] persuaded a publisher in the ] to pay ], and Darwin imported and distributed Gray's pamphlet ''Natural Selection is not inconsistent with ]''.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 492, 502}}.<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Miles|2001}}.</ref> In Britain, friends including Hooker<ref>{{Harvnb|Scott|2006}}.</ref> and ]<ref name=Bartholomew1976>{{Harvnb|Bartholomew|1976}}</ref> took part in the scientific debates which Huxley pugnaciously led to overturn the dominance of clergymen and aristocratic amateurs under Owen in favour of a new generation of professional scientists. Owen made the mistake of (wrongly) claiming certain anatomical differences between ape and ]s, and accusing Huxley of advocating "Ape Origin of Man". Huxley gladly did just that, and his campaign over two years was devastatingly successful in ousting Owen and the "old guard".<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 503–505}}.</ref> Darwin's friends formed '']'' and helped to gain him the honour of the ]'s Copley Medal in 1864.<ref name=Bartholomew1976 />
			The first review asked, "If a monkey has become a man – what may not a man become?" It said this should be left to theologians as being too dangerous for ordinary readers.<ref>{{harvnb|Browne|2002|p=87}}<br />{{harvnb|Leifchild|1859}}</ref> Among early favourable responses, Huxley's reviews swiped at ], leader of the scientific establishment which Huxley was trying to overthrow.<ref>{{Harvnb|Desmond|Moore|1991|pp=477–491}}</ref>
	Broader public interest had already been stimulated by '']'', and the ''Origin of Species'' was translated into many languages and went through numerous reprints, becoming a staple scientific text accessible both to a newly curious middle class and to "working men" who flocked to Huxley's lectures.<ref>{{Harvnb|Huxley|1863}}</ref> Darwin's theory also resonated with various movements at the time{{Ref_label|C|III|none}} and became a key fixture of popular culture.{{Ref_label|D|IV|none}}
			In April, Owen's review attacked Darwin's friends and condescendingly dismissed his ideas, angering Darwin,<ref>{{harvnb|Browne|2002|pp=110–112}}</ref> but Owen and others began to promote ideas of supernaturally guided evolution. ] drew attention to his 1831 book which had a brief appendix suggesting a concept of natural selection leading to new species, but he had not developed the idea.<ref>{{harvnb|Bowler|2003|pp=158, 186}}</ref>
			The ]'s response was mixed. Darwin's old Cambridge tutors Sedgwick and Henslow dismissed the ideas, but ] interpreted natural selection as an instrument of God's design, with the cleric ] seeing it as "just as noble a conception of Deity".<ref name="Darwin Correspondence Project-2007">{{cite web|url=http://www.darwinproject.ac.uk/content/view/110/104/|title=Darwin and design: historical essay|year=2007|publisher=Darwin Correspondence Project|access-date=17 September 2008 |archive-url=https://web.archive.org/web/20090615191012/http://www.darwinproject.ac.uk/content/view/110/104/ |archive-date=15 June 2009}}</ref> In 1860, the publication of '']'' by seven liberal Anglican theologians diverted ] attention from Darwin. Its ideas, including ], were attacked by church authorities as ]. In it, ] argued that ]s broke God's laws, so belief in them was ], and praised "Mr Darwin's masterly volume &#91;supporting&#93; the grand principle of the self-evolving powers of nature".<ref>{{Harvnb|Desmond|Moore|1991|pp=487–488, 500}}</ref>
			Asa Gray discussed ] with Darwin, who imported and distributed Gray's pamphlet on ], ''Natural Selection is not inconsistent with natural theology''.<ref name="Darwin Correspondence Project-2007" /><ref name="Miles 2001">{{Harvnb|Miles|2001}}</ref> The most famous confrontation was at the public ] during a meeting of the ], where the ] ], though not opposed to transmutation of species, argued against Darwin's explanation and human descent from apes. Joseph Hooker argued strongly for Darwin, and Thomas Huxley's legendary retort, that he would rather be descended from an ape than a man who misused his gifts, came to symbolise a triumph of science over religion.<ref name="Darwin Correspondence Project-2007" /><ref>{{harvnb|Bowler|2003|p=185}}</ref>
			Even Darwin's close friends Gray, Hooker, Huxley and Lyell still expressed various reservations but gave strong support, as did many others, particularly younger naturalists. Gray and Lyell sought reconciliation with faith, while Huxley portrayed a polarisation between religion and science. He campaigned pugnaciously against the authority of the clergy in education,<ref name="Darwin Correspondence Project-2007" /> aiming to overturn the dominance of clergymen and aristocratic amateurs under Owen in favour of a new generation of professional scientists. Owen's claim that brain anatomy proved humans to be a separate ] from apes was shown to be false by Huxley in a long-running dispute parodied by Kingsley as the "]", and discredited Owen.<ref>{{Harvnb|Browne|2002|pp=156–159}}</ref>
			In response to objections that the ] was unexplained, Darwin pointed to acceptance of ] even though the cause of gravity was unknown.<ref name="National University of Singapore News-2022">{{cite web | title=Science ahead of its time: Secret of 157-year old Darwin manuscript | website=] News | date=24 November 2022 | url=https://news.nus.edu.sg/secret-of-157-year-old-darwin-manuscript/ | access-date=25 November 2022 | archive-date=25 November 2022 | archive-url=https://web.archive.org/web/20221125100051/https://news.nus.edu.sg/secret-of-157-year-old-darwin-manuscript/ | url-status=live }}</ref> Despite criticisms and reservations related to this topic, he nevertheless proposed a prescient idea in an 1871 letter to Hooker in which he suggested the origin of life may have occurred in a "]".<ref>{{Cite journal |last1=Peretó |first1=Juli |last2=Bada |first2=Jeffrey L. |last3=Lazcano |first3=Antonio |date=1 October 2009 |title=Charles Darwin and the Origin of Life |url=https://doi.org/10.1007/s11084-009-9172-7 |journal=Origins of Life and Evolution of Biospheres |language=en |volume=39 |issue=5 |pages=395–406 |doi=10.1007/s11084-009-9172-7 |issn=1573-0875 |pmc=2745620 |pmid=19633921 |access-date=8 December 2023 |archive-date=18 December 2023 |archive-url=https://web.archive.org/web/20231218114111/https://link.springer.com/article/10.1007/s11084-009-9172-7 |url-status=live }}</ref>
			] became a movement covering a wide range of evolutionary ideas. In 1863, Lyell's '']'' popularised prehistory, though his caution on evolution disappointed Darwin. Weeks later Huxley's '']'' showed that anatomically, humans are apes, then '']'' by ] provided empirical evidence of natural selection.<ref name="Browne 2002">{{harvnb|Browne|2002|pp=217–226}}</ref> Lobbying brought Darwin Britain's highest scientific honour, the Royal Society's ], awarded on 3 November 1864.<ref>{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-4652.html|title=Darwin Correspondence Project&nbsp;– Letter 4652&nbsp;– Falconer, Hugh to Darwin, C. R., 3 Nov (1864)|access-date=1 December 2008|url-status=live|archive-url=https://web.archive.org/web/20081205084616/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-4652.html|archive-date=5 December 2008}}</ref> That day, Huxley held the first meeting of what became the influential "]" devoted to "science, pure and free, untrammelled by religious dogmas".<ref name="Darwin Correspondence Project-7">{{cite web|url=http://www.darwinproject.ac.uk/darwinletters/calendar/entry-4807.html#mark-4807.f8|title=Darwin Correspondence Project&nbsp;– Letter 4807&nbsp;– Hooker, J. D. to Darwin, C. R., (7–8 Apr 1865)|access-date=1 December 2008|url-status=live|archive-url=https://web.archive.org/web/20081205084621/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-4807.html#mark-4807.f8|archive-date=5 December 2008}}</ref> By the end of the decade, most scientists agreed that evolution occurred, but only a minority supported Darwin's view that the chief mechanism was natural selection.<ref>{{harvnb|Bowler|2003|p=196}}</ref>
			The ''Origin of Species'' was translated into many languages, becoming a staple scientific text attracting thoughtful attention from all walks of life, including the "working men" who flocked to Huxley's lectures.<ref>{{harvnb|Desmond|Moore|1991|pp=507–508}}<br />{{Harvnb|Browne|2002|pp=128–129, 138}}</ref> Darwin's theory resonated with various movements at the time{{Ref label|E|V|none}} and became a key fixture of popular culture.{{Ref label|F|VI|none}} Cartoonists parodied animal ancestry in an old tradition of showing humans with animal traits, and in Britain, these droll images served to popularise Darwin's theory in an unthreatening way. While ill in 1862, Darwin began growing a beard, and when he reappeared in public in 1866, caricatures of him as an ] helped to identify all forms of evolutionism with Darwinism.<ref name="Browne 2002-2">{{harvnb|Browne|2002|pp=373–379}}</ref>
			], America's first palaeontologist, was the first to provide solid fossil evidence to support Darwin's theory of evolution by unearthing the ancestors of the modern horse.<ref>Plate, Robert. ''The Dinosaur Hunters: Othniel C. Marsh and Edward D. Cope,'' pp. 69, 203-5, David McKay Company, Inc., New York, 1964.</ref> In 1877, Marsh delivered a very influential speech before the annual meeting of the American Association for the Advancement of Science, providing a demonstrative argument for evolution. For the first time, Marsh traced the evolution of vertebrates from fish all the way through humans. Sparing no detail, he listed a wealth of fossil examples of past life forms. The significance of this speech was immediately recognized by the scientific community, and it was printed in its entirety in several scientific journals.<ref>McCarren, Mark J. ''The Scientific Contributions of Othniel Charles Marsh,'' pp. 37-9, Peabody Museum of Natural History, Yale University, New Haven, Connecticut, 1993. {{ISBN|0-912532-32-7}}</ref><ref>Plate, Robert. ''The Dinosaur Hunters: Othniel C. Marsh and Edward D. Cope,'' pp. 188-9, David McKay Company, Inc., New York, 1964.</ref>
	===''Descent of Man'', sexual selection, and botany===		===''Descent of Man'', sexual selection, and botany===
	:''More detailed articles cover Darwin's life ], ] and ]''		{{further|Darwin from Orchids to Variation|Darwin from Descent of Man to Emotions|Darwin from Insectivorous Plants to Worms|label 1=Orchids to Variation|label 2=Descent of Man to Emotions|label 3=Insectivorous Plants to Worms}}
	]'s portrait of Darwin]]
			]
	Despite repeated bouts of illness during the last twenty-two years of his life, Darwin pressed on with his work. He had published an abstract of his theory, but more controversial aspects of his "big book" were still incomplete, including explicit evidence of humankind's descent from earlier animals, and exploration of possible causes underlying the development of society and of human mental abilities. He had yet to explain features with no obvious utility other than decorative beauty. His experiments, research and writing continued.
			Despite repeated bouts of illness during the last twenty-two years of his life,{{sfn|Desmond|Moore|Browne|2007|pp=73–75}} Darwin's work continued. Having published ''On the Origin of Species'' as an ] of his theory, he pressed on with experiments, research, and writing of his "]". He covered human descent from earlier animals, including the evolution of society and of mental abilities, as well as explaining decorative beauty in ] and diversifying into innovative plant studies.{{sfn|Desmond|Moore|Browne|2007|pp=78–83, 86–90}}
	When Darwin's daughter fell ill, he set aside his experiments with seedlings and domestic animals to accompany her to a seaside resort where he became interested in wild ]s. This developed into an innovative study of how their beautiful flowers served to control insect ] and ensure ]. As with the barnacles, homologous parts served different functions in different species. Back at home, he lay on his sickbed in a room filled with experiments on ]. A reverent Ernst Haeckel who had spread the gospel of Darwinismus in Germany visited him.<ref> Cambridge University Press. Retrieved on ].</ref> Wallace remained supportive, though he increasingly turned to ].<ref>{{harvnb|Smith|1999}}.</ref>
			Enquiries about insect ] led in 1861 to novel studies of wild ]s, showing adaptation of their flowers to ] to each species and ensure ]. In 1862 '']'' gave his first detailed demonstration of the power of natural selection to explain complex ecological relationships, making testable predictions. As his health declined, he lay on his sickbed in a room filled with inventive experiments to trace the movements of ].<ref>{{harvnb|van Wyhe|2008b|pp=50–55}}</ref> Admiring visitors included ], a zealous proponent of Darwinism incorporating Lamarckism and ]'s idealism.<ref>{{cite web |url=https://www.darwinproject.ac.uk/letters/darwins-life-letters/darwin-letters1866-survival-fittest |title=The correspondence of Charles Darwin, volume 14: 1866 |access-date=6 March 2009 |archive-url=https://web.archive.org/web/20100605110511/http://www.darwinproject.ac.uk/correspondence-volume-14 |archive-date=5 June 2010 }} Cambridge University Press. Retrieved 25 June 2012</ref> Wallace remained supportive, though he increasingly turned to ].<ref>{{harvnb|Smith|1999}}.</ref>
	''Variation of Plants and Animals Under Domestication'', the first part of Darwin's planned "big book" (expanding on his "abstract" published as ''The Origin of Species''), grew to two huge volumes, forcing him to leave out ] and ], and sold briskly despite its size.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 550}}.</ref> A further book of evidence, dealing with natural selection in the same style, was largely written, but remained unpublished until transcribed in 1975.<ref>{{Harvnb|Freeman|1977|pp=122–7}}</ref>
			Darwin's book '']'' (1868) was the first part of his planned "big book", and included his unsuccessful hypothesis of ] attempting to explain ]. It sold briskly at first, despite its size, and was translated into many languages. He wrote most of a second part, on natural selection, but it remained unpublished in his lifetime.<ref>{{Harvnb|Freeman|1977|p=}}</ref>
	] ] for 1882, published shortly before Darwin's death, depicts him amidst evolution from chaos to Victorian gentleman with the title ''Man Is But A Worm''.]]
	The question of human evolution had been taken up by his supporters (and detractors) shortly after the publication of '']'',<ref>See list of books at Retrieved on ].</ref> but Darwin's own contribution to the subject came more than ten years later with the two-volume '']'' published in 1871. In the second volume, Darwin introduced in full his concept of sexual selection to explain the evolution of human ], the differences between the human sexes, and the differentiation of human ]s, as well as the beautiful (and seemingly non-adaptive) plumage of birds.<ref name=DescentOfMan>{{Harvnb|Darwin|1871}}<br />{{Harvnb|Moore & Desmond|2004|Ref=CITEREFMooreDesmond2004}}</ref> A year later Darwin published his last major work, '']'', which focused on the evolution of human psychology and its continuity with the ]. He developed his ideas that the human ] and cultures were developed by natural and sexual selection,<ref>{{Harvnb|Darwin|1872}}</ref> an approach which has been revived in the last three decades with the emergence of ].<ref>{{Harvnb|Ghiselin|1973}}</ref> As he concluded in ''Descent of Man'', Darwin felt that, despite all of humankind's "noble qualities" and "exalted powers": "Man still bears in his bodily frame the indelible stamp of his lowly origin."<ref>{{Harvnb|Darwin|1871|p=}}</ref>
			Lyell had already popularised human prehistory, and Huxley had shown that anatomically humans are apes.<ref name="Browne 2002"/> With ''The Descent of Man, and Selection in Relation to Sex'' published in 1871, Darwin set out evidence from numerous sources that humans are animals, showing continuity of physical and mental attributes, and presented sexual selection to explain impractical animal features such as the ]'s plumage as well as human evolution of culture, differences between sexes, and physical and cultural racial classification, while emphasising that humans are all one species.<ref>{{Harvnb|Darwin|1871|pp=}}<br />{{Harvnb|Browne|2002|pp=339–343}}</ref> According to an editorial in Nature journal: "Although Charles Darwin opposed slavery and proposed that humans have a common ancestor, he also advocated a hierarchy of races, with white people higher than others."<ref>{{Cite journal |last1=Nobles |first1=Melissa |last2=Womack |first2=Chad |last3=Wonkam |first3=Ambroise |last4=Wathuti |first4=Elizabeth |date=8 June 2022 |title=Science must overcome its racist legacy: Nature's guest editors speak |journal=Nature |language=en |volume=606 |issue=7913 |pages=225–227 |doi=10.1038/d41586-022-01527-z |pmid=35676434 |bibcode=2022Natur.606..225N |s2cid=249520597 |quote="In The Descent of Man, Darwin describes what he calls the gradations between the highest men of the highest races and the lowest savages*. He uses the word 'savages' to describe Black and Indigenous people." (*see Darwin, C. R. The Descent of Man, and Selection in Relation to Sex (John Murray, 1871))|doi-access=free }}</ref>
	His evolution-related experiments and investigations culminated in books on the movement of climbing plants, ], the effects of ] and ] of plants, different forms of flowers on plants of the same species, and '']''. In his last book, he returned to the effect ]s have on soil formation.
			{{Multiple image|total_width=400
	He died in Downe, ], England, on ] ]. He had expected to be buried in St Mary's churchyard at Downe, but at the request of Darwin's colleagues, ] (President of the ]) arranged for Darwin to be given a ] and buried in ], close to ] and ].<ref>{{Harvnb|Browne|2002|pp=495–497}}.</ref>
			|image1=Darwin letter.jpg
			|alt1=handwritten letter from Charles Darwin to John Burdon-Sanderson dated 9 October 1874
			|caption1=Letter of enquiry from Charles Darwin to the physiologist ]
			|image2=Man is But a Worm.jpg
			|alt2=Darwin's figure is shown seated, dressed in a toga, in a circular frame labelled "TIME'S METER" around which a succession of figures spiral, starting with an earthworm emerging from the broken letters "CHAOS" then worms with head and limbs, followed by monkeys, apes, primitive men, a loin cloth clad hunter with a club, and a gentleman who tips his top hat to Darwin
			|caption2='']''{{'}}s ] for 1882, published shortly before Darwin's death, depicts him amidst evolution from chaos to Victorian gentleman with the title ''Man Is But A Worm''
			}}
			His research using images was expanded in his 1872 book '']'', one of the first books to feature printed photographs, which discussed the ] and its continuity with the ]. Both books proved very popular, and Darwin was impressed by the general assent with which his views had been received, remarking that "everybody is talking about it without being shocked."<ref>{{Harvnb|Browne|2002|pp=359–369}}<br />{{harvnb|Darwin|1887|p=}}</ref> His conclusion was "that man with all his noble qualities, with sympathy which feels for the most debased, with benevolence which extends not only to other men but to the humblest living creature, with his god-like intellect which has penetrated into the movements and constitution of the solar system{{snd}}with all these exalted powers{{snd}}Man still bears in his bodily frame the indelible stamp of his lowly origin."<ref>{{Harvnb|Darwin|1871|p=}}</ref>
	==Darwin's children==
	{| class="toccolours" style="float: right; margin-left: 1em; margin-right: 1em; font-size: 85%; background:#e0e0ee; color:black; width:32em; max-width: 50%;" cellspacing="5"
			His evolution-related experiments and investigations led to books on ''], ]'', different forms of flowers on plants of the same species, and '']''. He continued to collect information and exchange views from scientific correspondents all over the world, including ], whom he encouraged to persevere in her scientific work.<ref> {{Webarchive|url=https://web.archive.org/web/20200212213901/https://www.cam.ac.uk/research/news/darwins-women |date=12 February 2020 }} at ]</ref> He was the first person to recognise the significance of carnivory in plants.<ref name="Hedrich-2021">{{cite journal |last1=Hedrich |first1=Rainer |last2=Fukushima |first2=Kenji |title=On the Origin of Carnivory: Molecular Physiology and Evolution of Plants on an Animal Diet |journal=Annual Review of Plant Biology |date=17 June 2021 |volume=72 |issue=1 |pages=133–153 |doi=10.1146/annurev-arplant-080620-010429 |pmid=33434053 |s2cid=231595236 |issn=1543-5008|doi-access=free }}</ref> His botanical work{{Ref label|I|IX|none}} was interpreted and popularised by various writers including ] and ], and helped transform plant science in the late 19th century and early 20th century.<ref name="Pain-2022">{{cite journal |last1=Pain |first1=Stephanie |title=How plants turned predator |journal=Knowable Magazine |date=2 March 2022 |doi=10.1146/knowable-030122-1 |doi-access=free |url=https://knowablemagazine.org/article/living-world/2022/how-plants-turned-predator |access-date=11 March 2022 |archive-date=8 March 2022 |archive-url=https://web.archive.org/web/20220308103952/https://knowablemagazine.org/article/living-world/2022/how-plants-turned-predator |url-status=live }}</ref><ref name="Endersby-2016">{{cite journal |last1=Endersby |first1=Jim |title=Deceived by orchids: sex, science, fiction and Darwin |journal=The British Journal for the History of Science |date=June 2016 |volume=49 |issue=2 |pages=205–229 |doi=10.1017/S0007087416000352 |pmid=27278105 |s2cid=23027055 |url=http://journals.cambridge.org/action/displayAbstract?fromPage=online&aid=10359900&fulltextType=RA&fileId=S0007087416000352 |access-date=17 March 2022 |language=en |issn=0007-0874}}</ref>
	|colspan="2"|<center>]]]</center>
	<Center>Darwin and his eldest son ] in 1842.</center>
			===Death and funeral===
	|-
			{{See also|Darwin from Insectivorous Plants to Worms#Death}}
	!<Center>Darwin's Children</CENTER>
	|-
			] and Charles Darwin in the nave of ], London.]]
	|William Erasmus Darwin ||(] ]&ndash;1914)
			In 1882, he was diagnosed with what was called "]" which then meant ] and disease of the heart. At the time of his death, the physicians diagnosed "anginal attacks", and "heart-failure"; there has since been scholarly speculation about his ].<ref name="Colp-2008">{{cite book |title=Darwin's Illness |pages=116–120 |first=Ralph |last=Colp |doi=10.5744/florida/9780813032313.003.0014 |chapter=The Final {{sic|nolink=y|reason=error in source|Illnes}} |year=2008 |isbn=978-0-8130-3231-3 }}</ref><ref name="Clayton-2010">{{cite journal |last=Clayton |first=Julie |date=24 June 2010 |title=Chagas disease 101 |journal=Nature |volume=465 |issue=n7301_supp |pages=S4–S5 |doi=10.1038/nature09220 |pmid=20571553 |bibcode=2010Natur.465S...3C |s2cid=205221512 |doi-access=free}}</ref>
			He died at Down House on 19 April 1882. His last words were to his family, telling Emma, "I am not the least afraid of death{{mdash}}Remember what a good wife you have been to me{{mdash}}Tell all my children to remember how good they have been to me". While she rested, he repeatedly told Henrietta and Francis, "It's almost worthwhile to be sick to be nursed by you".<ref>{{cite web |url=http://darwin-online.org.uk/content/frameset?viewtype=side&itemID=CUL-DAR210.9&pageseq=16 |title= CUL-DAR210.9|author=Darwin, Emma|author-link=Emma Darwin |year=1882 |access-date=8 January 2009 |url-status=live |archive-url=https://web.archive.org/web/20090628080442/http://darwin-online.org.uk/content/frameset?viewtype=side&itemID=CUL-DAR210.9&pageseq=16 |archive-date=28 June 2009}}</ref>
			He had expected to be buried in St Mary's churchyard at ], but at the request of Darwin's colleagues, after public and parliamentary petitioning, ] (President of the Royal Society) arranged for Darwin to be honoured by ], close to John Herschel and ]. The funeral, held on Wednesday 26 April, was attended by thousands of people, including family, friends, scientists, philosophers and dignitaries.<ref>{{Harvnb|Desmond|Moore|1991|pp=664–677}}</ref><ref name="Westminster Abbey-2016">{{cite web | title=Westminster Abbey » Charles Darwin | website=Westminster Abbey | date=2 January 2016 | url=http://www.westminster-abbey.org/our-history/people/charles-darwin | access-date=2 January 2016 | url-status=live | archive-url=https://web.archive.org/web/20160304200905/http://www.westminster-abbey.org/our-history/people/charles-darwin | archive-date=4 March 2016 | df=dmy-all }}<br />{{Harvnb|Leff|2000|loc=}}</ref>
			==Children==
			{{further|Darwin–Wedgwood family#Charles Darwin}}
			{|class=toccolours style=float:right;clear:right;font-size:small;margin-left:1em;
			|]||style=text-align:right;|27 December 1839 –||8 September 1914
	|-		|-
	|] || (] ]&ndash;] ])		|]||style=text-align:right;|2 March 1841 –||23 April 1851
	|-		|-
	|Mary Eleanor Darwin || (] ]&ndash;] ])		|Mary Eleanor Darwin||style=text-align:right;|23 September 1842 –||16 October 1842
	|-		|-
	|] || (] ]&ndash;1929)		|]||style=text-align:right;|{{nobr|25 September 1843 –}}||17 December 1927
	|-		|-
	|] || (] ]&ndash;] ])		|]||style=text-align:right;|9 July 1845 –||7 December 1912
	|-		|-
			|Elizabeth Darwin||style=text-align:right;|8 July 1847 –||8 June 1926
	|] || (] ]&ndash;1926)
	|-		|-
	|] || (] ]&ndash;] ])		|]||style=text-align:right;|16 August 1848 –||{{nobr|19 September 1925}}
	|-		|-
	|] || (] ]&ndash;] ])		|]||style=text-align:right;|15 January 1850 –||26 March 1943
	|-		|-
	|] || (] ]&ndash;] ])		|]||style=text-align:right;|13 May 1851 –||29 September 1928
	|-		|-
	|] || (] ]&ndash;] ])		|Charles Waring Darwin||style=text-align:right;|6 December 1856 –||28 June 1858
	|}		|}
	The Darwins had ten children: two died in infancy, and Annie's death at the age of ten had a devastating effect on her parents. Charles was a devoted father and uncommonly attentive to his children.<ref name=whowas/> Whenever they fell ill he feared that they might have inherited weaknesses from ] due to the close family ties he shared with his wife and cousin, ]. He examined this topic in his writings, contrasting it with the advantages of crossing amongst many organisms.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p=447}}.</ref> Despite his fears, most of the surviving children went on to have distinguished careers as notable members of the prominent ].<ref> ''AboutDarwin.com''. Retrieved on ].</ref>
			The Darwins had ten children: two died in infancy, and Annie's death at the age of ten had a devastating effect on her parents. Charles was a devoted father and uncommonly attentive to his children.<ref name="Leff 2000" /> Whenever they fell ill, he feared that they might have inherited weaknesses from ] due to the close family ties he shared with his ], Emma Wedgwood. He examined inbreeding in his writings, contrasting it with the advantages of ] in many species.{{sfn|Desmond|Moore|1991|p=447}}
	Of his surviving children, ], ] and ] became Fellows of the Royal Society, distinguished as ],<ref>{{MacTutor Biography|id=Darwin}}</ref> ] and ], respectively.<ref>{{cite web | title=Royal Society Fellows' Directory | url=http://www.royalsoc.ac.uk/downloaddoc.asp?id=3120 | accessdate = 2006-12-15 | format=PDF}}</ref> His son ], on the other hand, went on to be a ], ], ], ] and mentor of the statistician and evolutionary biologist ].<ref>Edwards, A. W. F. 2004. Darwin, Leonard (1850–1943). In: ''Oxford Dictionary of National Biography'', Oxford University Press.</ref>
			]
	==Religious views==
	{{details|Charles Darwin's views on religion}}
			Charles Waring Darwin, born in December 1856, was the tenth and last of the children. Emma Darwin was aged 48 at the time of the birth, and the child was mentally subnormal and never learnt to walk or talk. He probably had ], which had not then been medically described. The evidence is a photograph by William Erasmus Darwin of the infant and his mother, showing a characteristic head shape, and the family's observations of the child.<ref>David P. Steensma (15 March 2005). "Down syndrome in Down House: trisomy 21, GATA1 mutations, and Charles Darwin". '']'' 105 (6) 2614–2616.</ref> Charles Waring died of scarlet fever on 28 June 1858,<ref>Freeman, R. B. (1984), ''Darwin Pedigrees'', London, p. 43.</ref> when Darwin wrote in his journal: "Poor dear Baby died."<ref>Darwin, C. R. ''Journal (1809–1881)'', p. 37.</ref> <!--The death kept Darwin from attending the first ] at the Linnean Society meeting on 1 July 1858.-->
	Though Charles Darwin's family background was ], and his father, grandfather and brother were ],<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 9, 12}}.</ref> at first he did not doubt the ] of the ].<ref>{{Harvnb|Darwin|1887|p=}}</ref> He attended a ] school, then at Cambridge studied ] theology to become a clergyman.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 12–15, 80–81}}.</ref> He was convinced by ]'s ] that design in nature proved the ],<ref>{{Harvnb|Darwin|1887|p=}}.</ref> but during the ''Beagle'' voyage he questioned, for example, why beautiful deep-ocean creatures had been created where no one could see them, or how the ] paralysing ]s as live food for its eggs could be reconciled with Paley's vision of beneficent design.<ref>{{Harvnb|Desmond|2004}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Lamoureux|2004|p=5}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Darwin|1887|p=}}.</ref> He was still quite ] and would quote the Bible as an authority on ], but did not trust the history in the ].<ref>{{Harvnb|Darwin|1958|p=}}.</ref>
			Of his surviving children, ], Francis and ] became ],<ref>{{cite web|title=List of Fellows of the Royal Society, 1660–2006, A–J|url=http://royalsociety.org/trackdoc.asp?id=4274&pId=1727|access-date=16 September 2009|format=PDF|archive-date=3 February 2017|url-status=dead|archive-url=https://web.archive.org/web/20170203000229/https://royalsociety.org/trackdoc.asp?id=4274&pId=1727}}</ref> distinguished as an astronomer,<ref>{{MacTutor Biography|id=Darwin}}</ref> botanist and civil engineer, respectively. All three were knighted.<ref>Berra, Tim M. ''Darwin and His Children: His Other Legacy,'' (Oxford: 2013, Oxford UP), 101, 129, 168. George became a knight commander of the Order of the Bath in 1905. Francis was knighted in 1912. Horace became a knight commander of the KBE in 1918.</ref> Another son, ], went on to be a soldier, politician, economist, ], and mentor of the statistician and evolutionary biologist Ronald Fisher.<ref>Edwards, A. W. F. 2004. Darwin, Leonard (1850–1943). In: ''Oxford Dictionary of National Biography'', Oxford University Press.</ref>
	], was the final step in pushing an already doubting Darwin away from the idea of a beneficent God.]]
	When investigating ] he knew that his naturalist friends thought this a bestial heresy undermining miraculous justifications for the social order, the kind of ] argument then being used by ]s and ] to attack the Church of England's privileged position as the ].<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 217–219, 221}}</ref> Though Darwin wrote of religion as a ] survival strategy, he still believed that God was the ultimate lawgiver.<ref>{{Harvnb|Moore|2006}}</ref> His belief dwindled, and with the death of his daughter ] in 1851, Darwin finally lost all faith in ]. He continued to help the local church with parish work, but on Sundays would go for a walk while his family attended church.<ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p= 387, 402}}</ref> He now thought it better to look at pain and suffering as the result of general laws rather than direct intervention by God.<ref>{{Harvnb|Darwin|1887|p=.}}</ref> When asked about his religious views, he wrote that he had never been an atheist in the sense of denying the existence of a God, and that generally "an ] would be the more correct description of my state of mind."<ref>{{Harvnb|Darwin|1887|p=.}}</ref>
			==Views and opinions==
	The "]", published in 1915, claimed that Darwin had reverted back to Christianity on his sickbed. The claims were refuted by Darwin's children and have been dismissed as false by historians.<ref>{{harvnb|Padian|n.d.}}.</ref> His daughter, Henrietta, who was at his deathbed, said that he did not convert to Christianity.<ref>{{Harvnb|Yates|2003}}</ref> His last words were, in fact, directed at Emma: ''"Remember what a good wife you have been."''<ref>{{Harvnb|Browne|2002|p=495}}.</ref>
	<br clear="both" />
			===Religious views===
	==Political interpretations==
			{{Further|Religious views of Charles Darwin}}
	]
			Darwin's family tradition was ] ], while his father and grandfather were ], and his baptism and boarding school were ].<ref name="Desmond-3" /> When going to Cambridge to become an Anglican clergyman, he did not "in the least doubt the strict and ] of every word in the Bible".<ref name="Desmond" /> He learned John Herschel's science which, like William Paley's natural theology, sought explanations in laws of nature rather than miracles and saw adaptation of species as evidence of design.<ref name="von Sydow 2005" /><ref name="Darwin 1958-5" /> On board HMS ''Beagle'', Darwin was quite ] and would quote the Bible as an authority on ].<ref name="Darwin 1958-4">{{Harvnb|Darwin|1958|pp=}}</ref> He looked for "centres of creation" to explain distribution,<ref name="Keynes 2001-2" /> and suggested that the very similar ]s found in Australia and England were evidence of a divine hand.<ref name="Darwin Online" />
	Darwin's theories and writings, combined with ]'s ] (the "]"), form the basis of all modern biology.<ref>{{Harvnb|Bowler|1989}}<br />&nbsp;&nbsp;&nbsp;{{harvnb|Dobzhansky|1973}}</ref> However, Darwin's fame and popularity led to his name being associated with ideas and movements which at times had only an indirect relation to his writings, and sometimes went directly against his express comments.
			] died; by then his faith in Christianity had dwindled, and he had stopped going to church.<ref name="van Wyhe 2008b">{{harvnb|van Wyhe|2008b|p=41}}</ref>]]
	===Eugenics===
	{{details|Eugenics}}
	Following Darwin's publication of the ''Origin'', his cousin, ], applied the concepts to human society, starting in 1865 with ideas to promote "hereditary improvement" which he elaborated at length in 1869.<ref>{{Harvnb|Galton|1865}} and {{Harvnb|Galton|1869}}</ref> In '']'' Darwin agreed that Galton had demonstrated the probability that "talent" and "genius" in humans was inherited, but dismissed the social changes Galton proposed as too ]n.<ref name=DoM5>{{Harvnb|Darwin|1871|loc=}}</ref> Neither Galton nor Darwin supported government intervention and thought that, at most, ] should be taken into consideration by people seeking potential mates.<ref>{{Harvnb|Galton|1869}} p. 1 and {{Harvnb|Darwin|1871|loc=}}</ref> In 1883, after Darwin's death, Galton began calling his social philosophy '']''.<ref>{{Harvnb|Galton|1883}} p 17, fn1.</ref> In the 20th century, eugenics movements gained popularity in a number of countries and became associated with reproduction control programmes such as ] laws,<ref>{{harvnb|Reilly|1991}}.</ref> then were stigmatised after their usage in the rhetoric of ] in its goals of genetic "purity".{{Ref_label|E|V|none}}
			Upon his return, he expressed a ] and questioned the basis for considering one religion more valid than another.<ref name="Darwin 1958-4" /> In the next few years, while intensively speculating on geology and the transmutation of species, he gave much thought to religion and openly discussed this with his wife Emma, whose beliefs similarly came from intensive study and questioning.<ref name="Darwin Correspondence Project" />
	===Social Darwinism ===
	{{details|Social Darwinism}}
	The ideas of ] and ] which applied ideas of evolution and "]" to societies, nations and businesses became popular in the late 19th and early 20th century, and were used to defend various, sometimes contradictory, ideological perspectives including ],<ref>{{Harvnb|Kotzin|2004}}</ref> ],<ref name=SocialDarwinismThinkQuest> ''ThinkQuest.org''. Retrieved on ].</ref> ] and ].<ref name=SocialDarwinismThinkQuest>{{cite web|url=http://library.thinkquest.org/C004367/eh4.shtml|title=''Social Darwinism'' at ThinkQuest.org|accessdate=2007-07-27}}</ref> The term "Social Darwinism" originated around the 1890s, but became popular as a derogatory term in the 1940s with ]'s critique of laissez-faire conservatism.<ref>{{Harvnb|Paul|2003}}</ref> The concepts predate Darwin's publication of the ''Origin'' in 1859:<ref name=SocialDarwinismThinkQuest /><ref>{{Harvnb|Desmond & Moore|1991|Ref=CITEREFDesmondMoore1991|p=477}}.<br />&nbsp;&nbsp;&nbsp;{{harvnb|Wilkins|1997}}.</ref> Malthus died in 1834<ref>{{Harvnb|Anonymous|1935}}</ref> and Spencer published his books on economics in 1851 and on evolution in 1855.<ref>{{Harvnb|Sweet|2004}}</ref> Darwin himself insisted that social policy should not simply be guided by concepts of struggle and selection in nature,<ref>{{Harvnb|Bannister|1989}}</ref> and that sympathy should be extended to all races and nations.<ref>{{Harvnb|Browne|1995|p= 244–246}}</ref>{{Ref_label|F|VI|none}}
			The ] of Paley and ] vindicated evils such as starvation as a result of a benevolent creator's laws, which had an overall good effect. To Darwin, natural selection produced the good of adaptation but removed the need for design,<ref>{{harvnb|von Sydow|2005|pp=8–14}}</ref> and he could not see the work of an omnipotent deity in all the pain and suffering, such as the ] paralysing ]s as live food for its eggs.<ref name="Miles 2001" /> Though he thought of religion as a ] survival strategy, Darwin was reluctant to give up the idea of ]. He was increasingly troubled by the ].<ref>{{harvnb|von Sydow|2005|pp=4–5, 12–14}}</ref><ref>{{Harvnb|Moore|2006}}</ref>
	==Commemoration ==
	]
	During Darwin's lifetime, many species and geographical features were given his name. An expanse of water adjoining the ] was named '']'' by ] after Darwin's prompt action, along with two or three of the men, saved them from being marooned on a nearby shore when a collapsing ] caused a large wave that would have swept away their boats,<ref>{{Harvnb|FitzRoy|1839|pp=}}</ref> and the nearby ] in the ] was named in celebration of Darwin's 25th birthday.<ref>. ''AboutDarwin.com'' Retrieved on ].</ref> When the ] was surveying ] in 1839, Darwin's friend ] sighted a natural harbour which the ship's captain ] named '']''.<ref name=NTDoPaI>{{cite web|url=http://web.archive.org/web/20060918153343/http://www.ipe.nt.gov.au/whatwedo/landinformation/place/origins/palmdarwin.html|title=Territory origins|
	accessdate=2006-12-15|publisher=Northern Territory Department of Planning and Infrastructure, Australia}}</ref> The settlement of ] ] was officially renamed ] in 1911. It became the capital city of Australia's ],<ref name=NTDoPaI /> which also boasts ]<ref>{{cite web|url=http://www.cdu.edu.au/|title=Charles Darwin University Homepage.|accessdate=2006-12-15}}</ref> and ].<ref> Northern Territory, Australia Government. Retrieved on ].</ref> ], founded in 1964, was named in honour of the Darwin family, partially because they owned some of the land it was on.<ref> Darwin College, Cambridge University website. Retrieved on ].</ref>
			Darwin remained close friends with the ] of Downe, ], and continued to play a leading part in the parish work of the church,<ref>{{cite web |url=https://www.darwinproject.ac.uk/commentary/religion/darwin-and-church |title=Darwin Correspondence Project&nbsp;– Darwin and the church: historical essay |access-date=26 November 2016 |url-status=live |archive-url=https://web.archive.org/web/20161128133709/https://www.darwinproject.ac.uk/commentary/religion/darwin-and-church |archive-date=28 November 2016 |date=5 June 2015 }}</ref> but from {{circa|1849}} would go for a walk on Sundays while his family attended church.<ref name="van Wyhe 2008b" /> He considered it "absurd to doubt that a man might be an ardent theist and an evolutionist"<ref name="Letter 12041 Archived 7 November 2009 at the Wayback Machine"> {{webarchive|url=https://web.archive.org/web/20091107174817/http://www.darwinproject.ac.uk/darwinletters/calendar/entry-12041.html |date=7 November 2009 }}&nbsp;– Darwin, C. R. to Fordyce, John, 7 May 1879</ref><ref name="Darwin"> {{Webarchive|url=https://web.archive.org/web/20170211082018/https://www.theguardian.com/commentisfree/belief/2009/sep/17/darwin-evolution-religion |date=11 February 2017 }} ] 17 September 2009</ref> and, though reticent about his religious views, in 1879 he wrote that "I have never been an atheist in the sense of denying the existence of a God.&nbsp;– I think that generally ... an agnostic would be the most correct description of my state of mind".<ref name="Darwin Correspondence Project"/><ref name="Letter 12041 Archived 7 November 2009 at the Wayback Machine"/>
	The 14 species of ]es he collected in the ] are affectionately named "]" in honour of his legacy.<ref>{{Harvnb|Rothman|2000}}.</ref> In 1992, Darwin was ranked #16 on ]'s ].<ref>{{Harvnb|Hart|2000|pp=82ff.}}</ref> Darwin came fourth in the '']'' poll sponsored by the ] and voted for by the public.<ref> ]. Retrieved on ].</ref> In 2000 Darwin's image appeared on the ] ], replacing ]. His impressive, luxuriant beard (which was reportedly difficult to forge) was said to be a contributory factor to the bank's choice.<ref> '']'' (] ]). Retrieved on ].</ref>
			The "]", published in 1915, claimed that Darwin had reverted to Christianity on his sickbed. The claims were repudiated by Darwin's children and have been dismissed as false by historians.<ref>{{harvnb|Moore|2005}}<br/>{{Harvnb|Yates|2003}}</ref>
	As a humorous celebration of evolution, the annual ] is bestowed on individuals who "improve our gene pool by removing themselves from it."<ref> ''Snopes.com'' (] ]). Retrieved on ].</ref>
			===Human society===
	Darwin has been the subject of many exhibitions, including the "Darwin" exhibition organised by the ] in ] in 2006 and shown in various cities in the US.<ref> ], Retrieved on ].</ref> Numerous biographies of Darwin have appeared, and the 1980 biographical novel '']'' by ] gives a closely researched fictional account of Darwin's life from the age of 22 onwards.
			Darwin's views on social and political issues reflected his time and social position. He grew up in a family of ] reformers who, like his uncle Josiah Wedgwood, supported ] and the emancipation of slaves. Darwin was passionately opposed to slavery, while seeing no problem with the working conditions of English factory workers or servants.{{sfn|Browne|1995|pp=196–197}}
			Taking taxidermy lessons in 1826 from the freed slave John Edmonstone, whom Darwin long recalled as "a very pleasant and intelligent man", reinforced his belief that black people shared the same feelings, and could be as intelligent as people of other races. He took the same attitude to native people he met on the ''Beagle'' voyage.{{sfn|Browne|1995|pp=66, 198, 240}} Though commonplace in Britain at the time, ] and ] noticed the contrast with slave-owning America. Around twenty years later, racism became a feature of British society,<ref name="Darwin 1958-6"/><ref name="Silliman-1810">{{cite book | last=Silliman | first=B. | title=A Journal of Travels in England, Holland and Scotland: And of Two Passages Over the Atlantic, in the Years 1805 and 1806 ... | publisher=D. & G. Bruce | year=1810 | url=https://books.google.com/books?id=z4pCAAAAYAAJ&pg=PA216 | access-date=29 August 2022 | pages=216–217 | quote=As there are no slaves in England, perhaps the English have not learned to regard negroes as a degraded class of men, as we do in the United States | archive-date=18 December 2023 | archive-url=https://web.archive.org/web/20231218114110/https://books.google.com/books?id=z4pCAAAAYAAJ&pg=PA216#v=onepage&q&f=false | url-status=live }}<br/>{{cite book | last=Bachman | first=J. | title=The Doctrine of the Unity of the Human Race Examined on the Principles of Science | publisher=C. Canning | series=American culture series | year=1850 | isbn=978-0-608-43507-7 | url=https://books.google.com/books?id=qKYdfBlX-GMC&pg=PA105 | access-date=29 August 2022 | page=105 | archive-date=18 September 2023 | archive-url=https://web.archive.org/web/20230918120545/https://books.google.com/books?id=qKYdfBlX-GMC&pg=PA105 | url-status=live }}</ref> but Darwin remained strongly against slavery, against "ranking the so-called races of man as distinct species", and against ill-treatment of native people.<ref>{{harvnb|Wilkins|2008|pp=408–413}}</ref>{{Ref label|G|VII|none}}
			Darwin's interaction with ] (Fuegians) such as Jemmy Button during the second voyage of HMS ''Beagle'' had a profound impact on his view of indigenous peoples. At his arrival in Tierra del Fuego he made a colourful description of "] savages".<ref name="Rozzi-2018" /> This view changed as he came to know Yaghan people more in detail. By studying the Yaghans, Darwin concluded that a number of basic emotions by different human groups were the same and that mental capabilities were roughly the same as for Europeans.<ref name="Rozzi-2018">{{cite journal |last1=Rozzi |first1=Ricardo|author-link=Ricardo Rozzi |date=2018 |title=Transformaciones del pensamiento de Darwin en cabo de hornos: Un legado para la ciencia y la etica ambiental|trans-title=Transformations of Darwin's thought in cape horn: A legacy for science and environmental ethics |language=es |journal=] |volume=46 |issue=1 |pages=267–277|doi=10.4067/S0718-22442018000100267 |doi-access=free }}</ref> While interested in Yaghan culture, Darwin failed to appreciate their deep ecological knowledge and elaborate cosmology until the 1850s when he inspected a dictionary of ] detailing 32,000 words.<ref name="Rozzi-2018" /> He saw that European colonisation would often lead to the extinction of native civilisations, and "tr to integrate colonialism into an evolutionary history of civilization analogous to natural history".<ref name="Barta-2005">{{cite journal |first=Tony|last=Barta|title=Mr Darwin's shooters: on natural selection and the naturalizing of genocide|journal=Patterns of Prejudice |volume=39|issue=2|pages=116–137 |doi=10.1080/00313220500106170 |date=2 June 2005 |s2cid=159807728}}</ref>
			] was that men's eminence over them was the outcome of sexual selection, a view disputed by ] in her 1875 book '']''.<ref name="Vandermassen, Griet-2004">{{cite journal |author=Vandermassen, Griet |title=Sexual Selection: A Tale of Male Bias and Feminist Denial |journal=European Journal of Women's Studies |year=2004 |volume=11 |issue=9 |doi=10.1177/1350506804039812 |pages=11–13 |citeseerx=10.1.1.550.3672 |s2cid=145221350 }}</ref>
			Darwin was intrigued by his ] ]'s argument, introduced in 1865, that ] of ] showed that moral and mental human traits could be inherited, and principles of animal breeding could apply to humans. In ''The Descent of Man'', Darwin noted that aiding the weak to survive and have families could lose the benefits of natural selection, but cautioned that withholding such aid would endanger the instinct of sympathy, "the noblest part of our nature", and factors such as education could be more important. When Galton suggested that publishing research could encourage intermarriage within a "caste" of "those who are naturally gifted", Darwin foresaw practical difficulties and thought it "the sole feasible, yet I fear ], plan of procedure in improving the human race", preferring to simply publicise the importance of inheritance and leave decisions to individuals.<ref>{{harvnb|Desmond|Moore|1991|pp=556–557, 572, 598}}<br />{{Harvnb|Darwin|1871|pp=, }}<br />{{cite web|url=http://www.galton.org/letters/darwin/correspondence.htm|title=Correspondence between Francis Galton and Charles Darwin|access-date=8 November 2008|url-status=live|archive-url=https://web.archive.org/web/20090102150434/http://galton.org/letters/darwin/correspondence.htm|archive-date=2 January 2009}}</ref> Francis Galton named this field of study "eugenics" in 1883,{{Ref label|H|VIII|1}} after Darwin's death, and his theories were cited to promote eugenic policies.<ref name="Barta-2005" />
			==Evolutionary social movements==
			{{further|Eugenics|Social effects of evolutionary theory|Degeneration theory}}
			]
			].]]
			Darwin's fame and popularity led to his name being associated with ideas and movements that, at times, had only an indirect relation to his writings, and sometimes went directly against his express comments.
			Thomas Malthus had argued that population growth beyond resources was ordained by God to get humans to ] and show restraint in getting families; this was used in the 1830s to justify ]s and ].<ref name="Wilkins 1997Moore 2006">{{harvnb|Wilkins|1997}}<br />{{Harvnb|Moore|2006}}</ref> Evolution was by then seen as having social implications, and ]'s 1851 book ''Social Statics'' based ideas of human freedom and individual liberties on his Lamarckian evolutionary theory.<ref>{{Harvnb|Sweet|2004}}</ref>
			Soon after the ''Origin'' was published in 1859, critics derided his description of a struggle for existence as a Malthusian justification for the English industrial capitalism of the time. The term ''Darwinism'' was used for the evolutionary ideas of others, including Spencer's "]" as free-market progress, and ]'s ] ideas of ]. Writers used natural selection to argue for various, often contradictory, ideologies such as laissez-faire dog-eat-dog capitalism, ] and ]. However, Darwin's holistic view of nature included "dependence of one being on another"; thus ], socialists, liberal social reformers and anarchists such as ] stressed the value of cooperation over struggle within a species.<ref>{{Harvnb|Paul|2003|pp=223–225}}</ref> Darwin himself insisted that social policy should not simply be guided by concepts of struggle and selection in nature.<ref>{{Harvnb|Bannister|1989}}</ref>
			After the 1880s, a eugenics movement developed on ideas of biological inheritance, and for scientific justification of their ideas appealed to some concepts of Darwinism. In Britain, most shared Darwin's cautious views on voluntary improvement and sought to encourage those with good traits in "positive eugenics". During the "Eclipse of Darwinism", a scientific foundation for eugenics was provided by ] ]. Negative eugenics to remove the "feebleminded" were popular in America, Canada and Australia, and ] introduced ] laws, followed by several other countries. Subsequently, ] brought the field into disrepute.{{Ref label|H|VIII|2}}
			The term "]" was used infrequently from around the 1890s, but became popular as a derogatory term in the 1940s when used by ] to attack the ] conservatism of those like ] who opposed reform and socialism. Since then, it has been used as a term of abuse by those opposed to what they think are the moral consequences of evolution.<ref>{{Harvnb|Paul|2003}}<br />{{Harvnb|Kotzin|2004}}</ref><ref name="Wilkins 1997Moore 2006"/>
	==Works==		==Works==
	{{details|List of works by Charles Darwin}}		{{further|Charles Darwin bibliography}}
	Darwin was a prolific author, and even without publication of his works on evolution would have had a considerable reputation as the author of '']'', as a geologist who had published extensively on ] and had solved the puzzle of the formation of ]s, and as a biologist who had published the definitive work on ]s. While '']'' dominates perceptions of his work, '']'' and '']'' had considerable impact, and his books on plants including '']'' were innovative studies of great importance, as was his final work on ''The Formation of Vegetable Mould Through the Action of Worms''.<ref>{{Harvnb|Balfour|1882}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|van Wyhe|2006}}<br />&nbsp;&nbsp;&nbsp;{{Harvnb|Anonymous|1882}}</ref>
			Darwin was a prolific writer. Even without the publication of his works on evolution, he would have had a considerable reputation as the author of ''The Voyage of the Beagle'', as a geologist who had published extensively on South America and had solved the puzzle of the formation of ]s, and as a biologist who had published the definitive work on barnacles. While ''On the Origin of Species'' dominates perceptions of his work, ''The Descent of Man'' and ''The Expression of the Emotions in Man and Animals'' had considerable impact, and his books on plants including ''The Power of Movement in Plants'' were innovative studies of great importance, as was his final work on '']''.<ref>{{Harvnb|Balfour|1882}}<br />{{Harvnb|van Wyhe|2008}}<br />{{Harvnb|Anonymous|1882}}</ref><ref>{{cite book|last=Brummitt|first=R. K.|author2=C. E. Powell|title=Authors of Plant Names |publisher=] |year=1992 |isbn=978-1-84246-085-6}}</ref>
	His writings are currently available at – the provides links to all of his publications, including alternative editions, contributions to books & periodicals, correspondence, life and letters, autobiography, as well as a complete bibliography and catalogue of his manuscripts. The works are free to read, but not ], and include publications still under ]. For unencumbered versions of his major works, see {{gutenberg author| id=Charles+Darwin | name=Charles Darwin}}.
			== Legacy and commemoration ==
			{{further|List of things named after Charles Darwin|List of taxa described by Charles Darwin|Commemoration of Charles Darwin}}
			{{Anchor|Legacy}}
			] building where he had studied]]
			As Alfred Russel Wallace put it, Darwin had "wrought a greater revolution in human thought within a quarter of a century than any man of our time{{snd}}or perhaps any time", having "given us a new conception of the world of life, and a theory which is itself a powerful instrument of research; has shown us how to combine into one consistent whole the facts accumulated by all the separate classes of workers, and has thereby revolutionised the whole study of nature".<ref name="Van Wyhe-2021">{{cite book | last=Van Wyhe | first=J. | title=Charles Darwin: The Man, His Great Voyage, and His Theory of Evolution | publisher=Rosen Publishing Group, Incorporated | series=Pioneers of Science | year=2021 | isbn=978-1-4994-7110-6 | url=https://books.google.com/books?id=JnZeEAAAQBAJ&pg=PA155 | access-date=23 May 2022 | pages=154–155 | archive-date=18 December 2023 | archive-url=https://web.archive.org/web/20231218114110/https://books.google.com/books?id=JnZeEAAAQBAJ&pg=PA155#v=onepage&q&f=false | url-status=live }}</ref> The paleoanthropologist ] states that "Darwin is rightly considered to be the preeminent evolutionary scientist of all time".{{sfn|Holliday|2021|p=5}}
			By around 1880, most scientists were convinced of evolution as descent with modification, though few agreed with Darwin that natural selection "has been the main but not the exclusive means of modification".<ref name="van Wyhe 2008Darwin 1872">{{Harvnb|van Wyhe|2008}}<br />{{harvnb|Darwin|1872|p=}}.</ref> During "]" scientists explored alternative mechanisms. Then ] incorporated ] in '']'',<ref>{{cite journal |url=http://www.genetics.org/content/154/4/1419.full |journal=Genetics |title=The Genetical Theory of Natural Selection |first=A. W. F. |last=Edwards |author-link=A. W. F. Edwards |date=1 April 2000 |volume=154 |issue=4 |pages=1419–1426 |doi=10.1093/genetics/154.4.1419 |pmid=10747041 |pmc=1461012 |url-status=live |archive-url=https://web.archive.org/web/20150924041631/http://www.genetics.org/content/154/4/1419.full |archive-date=24 September 2015 }}</ref> leading to ] and the ], which continues to develop.<ref name="Bowler 2003" /> Scientific discoveries have confirmed and validated Darwin's key insights.<ref name="Van Wyhe-2021" />
			Geographical features given his name include ]<ref>{{Harvnb|FitzRoy|1839|pp=}}.</ref> and ],<ref>{{harvnb|Leff|2000|loc=}}</ref> both named while he was on ], and ], named by his former shipmates on ], which eventually became the location of ], the capital city of Australia's ].<ref name="Northern Territory Department of Planning and Infrastructure, Australia">{{cite web|url=http://www.ipe.nt.gov.au/whatwedo/landinformation/place/origins/palmdarwin.html|archive-url=https://web.archive.org/web/20060918153343/http://www.ipe.nt.gov.au/whatwedo/landinformation/place/origins/palmdarwin.html|archive-date=18 September 2006|title=Territory origins| access-date=15 December 2006|publisher=Northern Territory Department of Planning and Infrastructure, Australia}}</ref> Darwin's name was given, ] or ], to numerous plants and animals, including many he had collected on the voyage.<ref>{{Cite book |last=Heard, Stephen B. |title=Charles Darwin's barnacle and David Bowie's spider : how scientific names celebrate adventurers, heroes, and even a few scoundrels |others=Damstra, Emily S. |date=2020 |isbn=978-0-300-25269-9 |publisher=Yale University Press |location=New Haven |oclc=1143645266}}</ref><ref>{{cite web |url=http://www.darwinfacts.com/ |title=Charles Darwin 200 years&nbsp;– Things you didn't know about Charles Darwin |access-date=23 May 2009 |url-status=live |archive-url=https://web.archive.org/web/20090528033253/http://darwinfacts.com/ |archive-date=28 May 2009 }}</ref> The Linnean Society of London began awards of the ] in 1908, to mark fifty years from the joint reading on 1 July 1858 of papers by Darwin and Wallace publishing their theory. Further awards were made in 1958 and 2008; since 2010, the awards have been annual.<ref name="The Linnean Society-2016">{{cite web |date=1 February 2016 |title=The Darwin-Wallace Medal |url=https://www.linnean.org/the-society/medals-awards-prizes-grants/the-darwin-wallace-medal |access-date=22 April 2022 |website=The Linnean Society |archive-date=29 March 2019 |archive-url=https://web.archive.org/web/20190329223140/https://www.linnean.org/the-society/medals-awards-prizes-grants/the-darwin-wallace-medal |url-status=live }}</ref> ], a postgraduate college at ] founded in 1964, is named after the Darwin family.<ref>{{cite web |title=Darwin College – Maps and directions – University of Kent |url=https://www.kent.ac.uk/maps/canterbury/canterbury-campus/building/darwin-college |url-status=live |archive-url=https://web.archive.org/web/20161031024930/https://www.kent.ac.uk/maps/canterbury/canterbury-campus/building/darwin-college |archive-date=31 October 2016 |access-date=30 October 2016 |website=www.kent.ac.uk}}</ref> From 2000 to 2017, UK £10 banknotes issued by the ] featured Darwin's portrait printed on the reverse,<ref name="BBC News-2000">{{cite web |date=7 November 2000 |title=How to join the noteworthy |url=http://news.bbc.co.uk/1/hi/uk/1009901.stm |access-date=24 April 2022 |website=BBC News |archive-date=30 June 2006 |archive-url=https://web.archive.org/web/20060630102007/http://news.bbc.co.uk/1/hi/uk/1009901.stm |url-status=live }}</ref><ref name="CBBC Newsround-2013">{{cite web |date=24 July 2013 |title=Author Jane Austen to feature on new £10 note |url=https://www.bbc.co.uk/newsround/23434198 |access-date=24 April 2022 |website=CBBC Newsround |archive-date=24 April 2022 |archive-url=https://web.archive.org/web/20220424113045/https://www.bbc.co.uk/newsround/23434198 |url-status=live }}</ref> along with a ] and ].<ref name="bankofengland.co.uk-2005">{{cite web |date=25 May 2005 |title=Bank of England – Banknotes – Current Banknotes – £10 |url=http://www.bankofengland.co.uk/banknotes/current/current_10.htm |url-status=dead |archive-url=https://web.archive.org/web/20050525230931/http://www.bankofengland.co.uk/banknotes/current/current_10.htm |archive-date=25 May 2005 |access-date=24 April 2022 |website=bankofengland.co.uk}}</ref>
	==See also==		==See also==
			{{Portal|Biology}}
	* ] &ndash; a species of ] named after Charles Darwin.
	* ] &ndash; a Galápagos tortoise, possibly collected by Darwin; died ] ] at an estimated age of 175.
			<!-- Please avoid repeating links above -->
			{{div col|colwidth=20em}}
			* '']''
			* '']'' (biographical drama film)
			* ]
			* ]
			* ]
			* ]
	* ]		* ]
			* ]
	* ] &ndash; an amateur evolutionary theorist and contemporary of Darwin.
			* ]
	* ] &ndash; the great-great grandson of Charles Darwin who wrote a book about him, his daughter, and human evolution
			* ]
			* ]
			* ]
			{{div col end}}
	==Notes==		==Notes==
			{{refbegin}}
	<div class="references-small">
			'''{{small|I}}.''' {{Note label|B|II|none}} ] was to become known after the voyage for ], but at this time he had considerable interest in Lyell's ideas, and they met before the voyage when Lyell asked for observations to be made in South America. FitzRoy's diary during the ascent of the River Santa Cruz in ] recorded his opinion that the plains were ]es, but on return, newly married to a very religious lady, he recanted these ideas.{{Harv|Browne|1995|pp=186, 414}}
	'''<small>I</small>.''' {{Note_label|A|I|none}} Darwin was eminent as a ], ], ], and ]; after working as a physician's assistant and two years as a ] was educated as a ]; and was trained in ].
			'''{{small|II}}.''' {{Note label|C|III|none}} In the section ] of Chapter XIII of ''On the Origin of Species'', Darwin commented on ] bone patterns between humans and other mammals, writing: "What can be more curious than that the hand of a man, formed for grasping, that of a mole for digging, the leg of the horse, the paddle of the porpoise, and the wing of the bat, should all be constructed on the same pattern, and should include the same bones, in the same relative positions?"<ref>{{harvnb|Darwin|1859|p=}}</ref> and in the concluding chapter: "The framework of bones being the same in the hand of a man, wing of a bat, fin of the porpoise, and leg of the horse … at once explain themselves on the theory of descent with slow and slight successive modifications."<ref>{{harvnb|Darwin|1859|p=}}</ref>
	'''<small>II</small>.''' {{Note_label|B|II|none}} <!--bit off topic and inaccurate refs – Darwin became famous for his 1839 book '']'' (before '']'' in 1859). See '''', Professor Fred L. Wilson, ], ].-->] was to become known after the voyage for biblical literalism, but at this time he had considerable interest in Lyell's ideas, and they met before the voyage when Lyell asked for observations to be made in South America. FitzRoy's diary during the ascent of the River Santa Cruz in ] recorded his opinion that the plains were ]es, but on return, newly married to a very religious lady, he recanted these ideas.<ref>{{Harvnb|Browne|1995|pp= 186, 414}}.</ref>
			'''{{small|III}}.''' {{Note label|D|IV|1}}{{Note label|D|IV|2}}{{Note label|D|IV|3}}
	'''<small>III</small>.''' {{Note_label|C|III|none}} See, for example, WILLA volume 4, by Deborah M. De Simone: "Gilman shared many basic educational ideas with the generation of thinkers who matured during the period of "intellectual chaos" caused by Darwin's Origin of the Species. Marked by the belief that individuals can direct human and social evolution, many progressives came to view education as the panacea for advancing social progress and for solving such problems as urbanisation, poverty, or immigration."
			In '']'' Darwin mentioned ] in his concluding remark that "In the distant future I see open fields for far more important researches. Psychology will be based on a new foundation, that of the necessary acquirement of each mental power and capacity by gradation. Light will be thrown on the origin of man and his history."<ref name="Darwin 1859"/>
			In "Chapter VI: Difficulties on Theory" he referred to ]: "I might have adduced for this same purpose the differences between the races of man, which are so strongly marked; I may add that some little light can apparently be thrown on the origin of these differences, chiefly through sexual selection of a particular kind, but without here entering on copious details my reasoning would appear frivolous."<ref name="Darwin 1859-2"/>
	'''<small>IV</small>.''' {{Note_label|D|IV|none}} See, for example, the song "A lady fair of lineage high" from ]'s '']'', which describes the descent of man (but not woman!) from apes.
			In '']'' of 1871, Darwin discussed the first passage:
	'''<small>V</small>.''' {{Note_label|E|V|none}} The Nazi eugenics policies are discussed in a number of sources. A few of the more definitive ones are Robert Proctor, ''Racial hygiene: Medicine under the Nazis'' (Cambridge, MA: Harvard University Press, 1988) and Dieter Kuntz, ed., ''Deadly medicine: creating the master race'' (Washington, D.C.: United States Holocaust Memorial Museum, 2004) (). On the development of the ] movement before National Socialism, see Paul Weindling, ''Health, race and German politics between national unification and Nazism, 1870–1945'' (New York: Cambridge University Press, 1989).
			"During many years I collected notes on the origin or descent of man, without any intention of publishing on the subject, but rather with the determination not to publish, as I thought that I should thus only add to the prejudices against my views. It seemed to me sufficient to indicate, in the first edition of my 'Origin of Species,' that by this work 'light would be thrown on the origin of man and his history;' and this implies that man must be included with other organic beings in any general conclusion respecting his manner of appearance on this earth."<ref>{{Harvnb|Darwin|1871|p=}}</ref> In a preface to the 1874 second edition, he added a reference to the second point: "it has been said by several critics, that when I found that many details of structure in man could not be explained through natural selection, I invented sexual selection; I gave, however, a tolerably clear sketch of this principle in the first edition of the 'Origin of Species,' and I there stated that it was applicable to man."<ref>{{Harvnb|Darwin|1874|p=}}</ref>
			'''{{small|IV}}.''' {{Note label|E|V|none}} See, for example, WILLA volume 4, '''' by Deborah M. De Simone: "Gilman shared many basic educational ideas with the generation of thinkers who matured during the period of "intellectual chaos" caused by Darwin's Origin of the Species. Marked by the belief that individuals can direct human and social evolution, many progressives came to view education as the panacea for advancing social progress and for solving such problems as urbanisation, poverty, or immigration."
	'''<small>VI</small>.''' {{Note_label|F|VI|none}} See {{Harvnb|Darwin|1887|p=}}:
	:]arly in the voyage at Bahia, in Brazil, ] defended and praised slavery, which I abominated, and told me that he had just visited a great slave-owner, who had called up many of his slaves and asked them whether they were happy, and whether they wished to be free, and all answered "No." I then asked him, perhaps with a sneer, whether he thought that the answer of slaves in the presence of their master was worth anything? This made him excessively angry, and he said that as I doubted his word we could not live any longer together.
			'''{{small|V}}.''' {{Note label|F|VI|none}} See, for example, the song "A lady fair of lineage high" from ]'s '']'', which describes the descent of man (but not woman!) from apes.
	See also {{Harvnb|Darwin|1845|pp= }} on the ]:
	:It seems yet wonderful to me, when I think over all his many good qualities, that he should have been of the same race, and doubtless partaken of the same character, with the miserable, degraded savages whom we first met here.
			'''{{small|VI}}.''' {{Note label|G|VII|none}} Darwin's belief that black people had the same essential humanity as Europeans, and had many mental similarities, was reinforced by the lessons he had from ] in 1826.<ref name="Darwin 1958-6" /> Early in the ''Beagle'' voyage, Darwin nearly lost his position on the ship when he criticised FitzRoy's defence and praise of slavery. {{Harv|Darwin|1958|p=}} He wrote home about "how steadily the general feeling, as shown at elections, has been rising against Slavery. What a proud thing for England if she is the first European nation which utterly abolishes it! I was told before leaving England that after living in slave countries all my opinions would be altered; the only alteration I am aware of is forming a much higher estimate of the negro character." {{harv|Darwin|1887|p=}} Regarding ], he "could not have believed how wide was the difference between savage and civilized man: it is greater than between a wild and domesticated animal, inasmuch as in man there is a greater power of improvement", but he knew and liked civilised Fuegians like ]: "It seems yet wonderful to me, when I think over all his many good qualities, that he should have been of the same race, and doubtless partaken of the same character, with the miserable, degraded savages whom we first met here." {{Harv|Darwin|1845|pp=}}
	</div>
			In the ''Descent of Man'', he mentioned the similarity of Fuegians' and Edmonstone's minds to Europeans' when arguing against "ranking the so-called races of man as distinct species".<ref>{{Harvnb|Darwin|1871|pp=, }}</ref>
			He rejected the ill-treatment of native people, and for example wrote of massacres of ]n men, women, and children, "Every one here is fully convinced that this is the most just war, because it is against barbarians. Who would believe in this age that such atrocities could be committed in a Christian civilized country?"{{harv|Darwin|1845|p=}}
			'''{{small|VII}}.''' {{Note label|H|VIII|1}}{{Note label|H|VIII|2}} ]s studied human heredity as ], while ] movements sought to manage society, with a focus on social class in the United Kingdom, and on disability and ethnicity in the United States, leading to geneticists seeing this movement as impractical ]. A shift from voluntary arrangements to "negative" eugenics included ] laws in the United States, copied by ] as the basis for ] based on virulent racism and "]".<br />({{cite news | url=http://www.stanford.edu/group/SHR/5-supp/text/thurtle.html | title=the creation of genetic identity | last=Thurtle | first=Phillip | date=17 December 1996 | issue=Supplement: Cultural and Technological Incubations of Fascism | volume=5 | periodical=SEHR | access-date=11 November 2008 |ref=none}} {{Cite news |url=http://www.genetics.org/cgi/content/full/154/4/1419#The_Eclipse_of_Darwinism| title=The Genetical Theory of Natural Selection | last=Edwards | first=A. W. F. |author-link=A. W. F. Edwards | date=1 April 2000 | issue=April 2000 | volume=154 | pages=1419–1426 | pmc=1461012 | pmid=10747041 | periodical=Genetics | access-date=11 November 2008 | ref=none}}{{cite web |url=http://scienceblogs.com/evolvingthoughts/2006/09/darwin_and_the_holocaust_3_eug_1.php |title=Evolving Thoughts: Darwin and the Holocaust 3: eugenics |last=Wilkins |first=John |access-date=11 November 2008 |url-status=dead |archive-url=https://web.archive.org/web/20081205154013/http://scienceblogs.com/evolvingthoughts/2006/09/darwin_and_the_holocaust_3_eug_1.php |archive-date=5 December 2008 |ref=none}})
			'''{{small|VIII}}.''' {{Note label|I|IX|none}} ] writes of his "theory that turned to these arcane botanical studies – producing more than one book that was solidly empirical, discreetly evolutionary, yet a 'horrid bore' – at least partly so that the clamorous controversialists, fighting about apes and angels and souls, would leave him... alone". ], "The Brilliant Plodder" (review of Ken Thompson, ''Darwin's Most Wonderful Plants: A Tour of His Botanical Legacy'', ], 255 pp.; Elizabeth Hennessy, ''On the Backs of Tortoises: Darwin, the Galápagos, and the Fate of an Evolutionary Eden'', ], 310 pp.; Bill Jenkins, ''Evolution Before Darwin: Theories of the Transmutation of Species in Edinburgh, 1804–1834'', ], 222 pp.), '']'', vol. LXVII, no. 7 (23 April 2020), pp.&nbsp;22–24. Quammen, quoted from p.&nbsp;24 of his review.
			{{refend}}
	==Citations==		==Citations==
			{{reflist|30em}}
	<div class="references-small" style="column-count:3;-moz-column-count:3;">
	<references />
	</div>
	==References==		==Bibliography==
	*{{Harvard reference
			{{refbegin}}
	| Surname = Anonymous
			* {{cite news
	| Year = 1882
			| last=Anonymous
	| Title = Obituary: Death Of Chas. Darwin
			| year=1882
	| Journal = ]
			| title=Obituary: Death Of Chas. Darwin
	| Issue = April 21, 1882
			| periodical=The New York Times
	| URL = http://www.nytimes.com/learning/general/onthisday/bday/0212.html }} Retrieved on ]
			| issue=21 April 1882
	*{{Harvard reference
			| url=https://www.nytimes.com/learning/general/onthisday/bday/0212.html
	| Surname = Anonymous
			| access-date=30 October 2008
	| Year = 1935
			| archive-date=15 October 2009
	| Title = Obituary: Thomas Robert Malthus: Died ] ]
			| archive-url=https://web.archive.org/web/20091015051211/http://nytimes.com/learning/general/onthisday/bday/0212.html
	| Journal = Journal of the Royal Statistical Society,
			| url-status=live
	| Issue = Vol. 98, No. 2
	| Pages = 376–379.
	| URL = http://links.jstor.org/sici?sici=0952-8385%281935%2998%3A2%3C376%3AOTRMD2%3E2.0.CO%3B2-6 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Anonymous
	| Year = 1960
	| Date = June 1960
	| Title = Darwin as a Traveller
	| Journal = The Geographical Journal
	| Issue = Vol. 126, No. 2.
	| Pages = 129–136
	| URL = http://links.jstor.org/sici?sici=0016-7398%28196006%29126%3A2%3C129%3ADAAT%3E2.0.CO%3B2-P }} Retrieved on ]
	*{{Harvard reference
	| Surname = Arrhenius
	| Given = O.
	| Year = 1921
	| Date = October 1921
	| Title = Influence of Soil Reaction on Earthworms
	| Journal = Ecology
	| Issue = Vol. 2, No. 4
	| Pages = 255–257
	| URL = http://links.jstor.org/sici?sici=0012-9658%28192110%292%3A4%3C255%3AIOSROE%3E2.0.CO%3B2-P }} Retrieved on ]
	*{{Harvard reference
	| Surname = Balfour
	| Given = J. B.
	| Year = 1882
	| Date = 11 May 1882
	| Title = ]
	| Journal = Transactions & Proceedings of the Botanical Society of Edinburgh
	| Issue = 14
	| Pages = 284–298
	| URL = }} Retrieved on ]
	*{{Harvard reference
	| Surname = Bannister
	| Given = Robert C.
	| Year = 1989
	| Title = Social Darwinism: Science and Myth in Anglo-American Social Thought.
	| Publisher = Philadelphia: Temple University Press
	| ID = ISBN 0-87722-566-4}}
	*{{Harvard reference
	| Surname1 = Bartholomew
	| Given1 = M. J.
	| Year = 1976
	| Title = The Award of the Copley Medal to Charles Darwin
	| URL = http://links.jstor.org/sici?sici=0035-9149(197601)30%3A2%3C209%3ATAOTCM%3E2.0.CO%3B2-6
	| Journal = Notes and Records of the Royal Society of London
	| Issue = Vol. 30, No. 2 (Jan. 1976)
	| Pages = 209–218 }} (Includes Lyell's speech at the award ceremony which was his first public affiliation with Darwin's theory.) Retrieved on ]
	*{{Harvard reference
	| Surname = Bowler
	| Given = Peter J.
	| Authorlink =
	| Year = 1989
	| Title = The Mendelian Revolution: The Emergence of Hereditarian Concepts in Modern Science and Society
	| Publisher = Baltimore: Johns Hopkins University Press
	| ID = ISBN 0-485-11375-9 }}
	*{{Harvard reference
	| Surname = Browne
	| Given = E. Janet
	| Authorlink = Janet Browne
	| Year = 1995
	| Title = Charles Darwin: vol. 1 Voyaging
	| Publisher = London: Jonathan Cape
	| ID = ISBN 1-84413-314-1}}
	*{{Harvard reference
	| Surname = Browne
	| Given = E. Janet
	| Authorlink = Janet Browne
	| Year = 2002
	| Title = Charles Darwin: vol. 2 The Power of Place
	| Publisher = London: Jonathan Cape
	| ID = ISBN 0-7126-6837-3}}
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1835
	| Title = Extracts from letters to Professor Henslow. Cambridge
	| Publisher =
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F1&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1839
	| Title = Journal and Remarks (])
	| Publisher = London: Henry Colburn
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F20&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Darwin
	| Given1 = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Chapter = Pencil Sketch of 1842
	| Editor = Darwin, Francis
	| Year = 1842
	| Date = 1842 (published 1909)
	| Title = The foundations of The origin of species: Two essays written in 1842 and 1844.
	| Publisher = Cambridge University Press
	| Location = Cambridge
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F1556&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1845
	| Title = Journal of Researches (])
	| Edition = Second
	| Publisher = London: John Murray
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F14&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1859
	| Title = On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life
	| Publisher = London: John Murray
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=1 }} (]) Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1868
	| Title = The variation of animals and plants under domestication
	| Publisher = London: John Murray
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F880.1&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1871
	| Title = The Descent of Man and Selection in Relation to Sex
	| Publisher = London: John Murray
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F937.1&viewtype=text&pageseq=1 }} (]) Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1872
	| Title = The Expression of the Emotions in Man and Animals
	| Publisher = London: John Murray
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F1142&viewtype=text&pageseq=1 }} Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1887
	| Editor = ]
	| Title = The life and letters of Charles Darwin, including an autobiographical chapter.
	| Publisher = London: John Murray
	| URL = http://www.gutenberg.org/catalog/world/readfile?fk_files=39003&pageno=1 }} (]) Retrieved on ]
	*{{Harvard reference
	| Surname = Darwin
	| Given = Charles
	<!-- NOT NEEDED ON THIS PAGE, BUT POSSIBLY ELSEWHERE | Authorlink = Charles Darwin -->
	| Year = 1958
	| Editor = ]
	| Title = The autobiography of Charles Darwin 1809–1882. With the original omissions restored. Edited and with appendix and notes by his grand-daughter Nora Barlow.
	| Publisher = London: Collins
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F1497&viewtype=text&pageseq=1
	}} (]) Retrieved on ]
	*{{Harvard reference
	| Surname1 = Desmond
	| Given1 = Adrian J.
	| Year = 2004
	| Chapter = Darwin
	| Title = Encyclopaedia Britannica 2004 DVD
	}}		}}
			* {{cite journal
	*{{Harvard reference
			| last=Balfour
	| Surname1 = Desmond
			| first=J. H.
	| Given1 = Adrian
			| author-link=John Hutton Balfour
	| Surname2 = Moore
			| date=11 May 1882
	| Given2 = James
			| title=Obituary Notice of Charles Robert Darwin
	| Authorlink2 = James Moore (biographer)
			| journal=]
	| Year = 1991
			| issue=14
	| Title = Darwin
			| pages=284–298| title-link=s:Transactions & Proceedings of the Botanical Society of Edinburgh/Obituary Notice of Charles Robert Darwin
	| Publisher = London: Michael Joseph, Penguin Group
	| ID = ISBN 0-7181-3430-3}}
	*{{Harvard reference
	| Surname1 = Dobzhansky
	| Given1 = Theodosius
	| Year = 1973
	| Date = March 1973
	| Title = Nothing in Biology Makes Sense Except in the Light of Evolution
	| URL = http://www.2think.org/dobzhansky.shtml
	| Journal = The American Biology Teacher
	| Volume = 35
	| Pages = 125–129 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Eldredge
	| Given1 = Niles
	| Year = 2006
	| Title = Confessions of a Darwinist
	| URL = http://www.vqronline.org/articles/2006/spring/eldredge-confessions-darwinist/#fn11
	| Journal = The Virginia Quarterly Review
	| Issue = Spring 2006
	| Pages = 32–53 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = FitzRoy
	| Given1 = Robert
	| Authorlink1 = Robert Fitzroy
	| Year = 1839
	| Title = Voyages of the Adventure and Beagle, Volume II
	| URL = http://darwin-online.org.uk/content/frameset?itemID=F10.2&viewtype=text&pageseq=1
	| Publisher = Henry Colburn
	| Location = Great Marlborough Street, London }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Freeman
	| Given1 = R. B.
	| Year = 1977
	| Title = The Works of Charles Darwin: An Annotated Bibliographical Handlist
	| URL = http://darwin-online.org.uk/content/frameset?itemID=A1&viewtype=text&pageseq=1
	| Publisher = Wm Dawson & Sons Ltd
	| Location = Cannon House Folkestone, Kent, England
	| Edition = Second }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Galton
	| Given1 = Francis
	| Authorlink1 = Francis Galton
	| Year = 1865
	| Title = Hereditary talent and character
	| URL = http://www.mugu.com/galton/essays/1860-1869/galton-1865-hereditary-talent.pdf
	| Journal = Macmillan's Magazine
	| Issue = 12
	| Pages = 157–166 and 318–327 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Galton
	| Given1 = Francis
	| Authorlink1 = Francis Galton
	| Year = 1869
	| Title = Hereditary genius: an inquiry into its laws and consequences
	| Publisher = Macmillan
	| Location = London
	| URL = http://www.mugu.com/galton/books/hereditary-genius/
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Galton
	| Given1 = Francis
	| Authorlink1 = Francis Galton
	| Year = 1883
	| Title = Inquiries into human faculty and its development
	| Publisher = Macmillan
	| Location = London
	| URL = http://www.mugu.com/galton/books/human-faculty/
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Ghiselin
	| Given1 = Michael T.
	| Year = 1973
	| Title = Darwin and Evolutionary Psychology
	| URL = http://www.robertgordon.net/papers/four.html
	| Journal = Science
	| Volume = 179
	| Pages = 964–968 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Gordon
	| Given1 = Robert
	| Surname2 = Thomas
	| Given2 = Deborah
	| Year = 1999
	| Date = ]-] ]
	| Title = Circumnavigating Darwin
	| URL = http://www.robertgordon.net/papers/four.html
	| Journal = Darwin Undisciplined Conference, Sydney.
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Hart
	| Given1 = Michael
	| Authorlink = Michael Hart
	| Year = 2000
	| Title = The 100: A Ranking of the Most Influential Persons in History
	| Publisher = Citadel
	| Location = New York
	}}		}}
			* {{cite book
	*{{Harvard reference
			| last=Bannister
	| Surname = Huxley
			| first=Robert C.
	| Given = Thomas
			| year=1989
	| Authorlink = Thomas Huxley
			| title=Social Darwinism: Science and Myth in Anglo-American Social Thought.
	| Year = 1863
			| location=Philadelphia
	| Title = Six Lectures to Working Men "On Our Knowledge of the Causes of the Phenomena of Organic Nature" (Republished in Volume II of his Collected Essays, ''Darwiniana'')
			| publisher=Temple University Press
	| URL = http://aleph0.clarku.edu/huxley/CE2/Phen.html }} Retrieved on ]
			| isbn=978-0-87722-566-9}}
	*{{Harvard reference
			* {{cite book
	| Surname = Huxley
			| last=Bowler
	| Given = Thomas
			| first=Peter J.
	| Authorlink = Thomas Huxley
	| Year = 1897		| year=2003
	| Title = Evolution and Ethics and Other Essays		| title=Evolution: The History of an Idea
			| edition=3rd
	| Publisher = New York, D. Appleton and Company
			| publisher=University of California Press
	| URL = http://aleph0.clarku.edu/huxley/CE9/index.html
			| isbn=978-0-520-23693-6
	}} Retrieved on ]
			| url-access=registration
	*{{Harvard reference
			| url=https://archive.org/details/evolutionhistory0000bowl_n7y8
	| Surname1 = Keynes
	| Given1 = Richard (ed.)
	| Year = 2000
	| Title = Charles Darwin's zoology notes & specimen lists from H.M.S. Beagle.
	| Publisher = Cambridge University Press
	| Location = Cambridge
	| Chapter = }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Kotzin
	| Given1 = Daniel
	| Year = 2004
	| Title = Point-Counterpoint: Social Darwinism
	| Publisher = Columbia American History Online
	| URL = http://caho-test.cc.columbia.edu/pcp/14008.html }}
	*{{Harvard reference
	| Surname1 = Lamoureux
	| Given1 = Denis O.
	| Year = 2004
	| Date = March 2004
	| Title = Theological Insights from Charles Darwin
	| URL = http://users.ox.ac.uk/~jrlucas/legend.html
	| Journal = Perspectives on Science and Christian Faith
	| Volume = 56
	| Issue = 1
	| Pages = 2–12
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Leff
	| Given1 = David
	| Year = 2000
	| Title = About Charles Darwin
	| URL = http://www.aboutdarwin.com/darwin/WhoWas.html
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Lucas
	| Given1 = J. R.
	| Year = 1979
	| Title = Wilberforce and Huxley: A Legendary Encounter
	| URL = http://users.ox.ac.uk/~jrlucas/legend.html
	| Journal = The Historical Journal
	| Volume = 22
	| Issue = 2
	| Pages = 313–330
	}} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Malthus
	| Given1 = Thomas Robert
	| Authorlink1 = Thomas Malthus
	| Year = 1826
	| URL = http://www.econlib.org/library/Malthus/malPlong.html
	| Title = An Essay on the Principle of Population: A View of its Past and Present Effects on Human Happiness; with an Inquiry into Our Prospects Respecting the Future Removal or Mitigation of the Evils which It Occasions
	| Edition = Sixth
	| Publisher = John Murray
	| Location = London }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Miles
	| Given1 = Sara Joan
	| Year = 2001
	| Title = Charles Darwin and Asa Gray Discuss Teleology and Design
	| URL = http://www.asa3.org/ASA/PSCF/2001/PSCF9-01Miles.html
	| Journal = Perspectives on Science and Christian Faith
	| Volume = 53
	| Pages = 196–201 }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Moore
	| Given1 = James
	| Authorlink1 = James Moore (biographer)
	| Surname2 = Desmond
	| Given2 = Adrian
	| Year = 2004
	| Title = "Introduction", in Darwin's The Descent of Man and Selection in Relation to Sex
	| Publisher = London: Penguin Classics
	}}
	*{{Harvard reference
	| Surname1 = Moore
	| Given1 = James
	| Authorlink1 = James Moore (biographer)
	| Year = 2006
	| Chapter = Evolution and Wonder - Understanding Charles Darwin
	| URL = http://speakingoffaith.publicradio.org/programs/darwin/transcript.shtml
	| Title = Speaking of Faith (Radio Program)
	| Publisher = American Public Media }} Retrieved on ]
	*{{Harvard reference
	| Surname1 = Owen
	| Given1 = Richard
	| Authorlink = Richard Owen
	| Editor = Darwin, C. R.
	| Year = 1840
	| Title = The zoology of the voyage of H.M.S. Beagle., Fossil Mammalia Part 1
	| Publisher = Smith Elder and Co
	| Location = London
	}}		}}
			* {{cite book
	*{{Harvard reference
			| last=Browne
	| Surname1 = Padian
	| Given1 = Kevin		| first=E. Janet
			| author-link=Janet Browne
	| Year = n.d.
			| year=1995
	| Title = The Darwin Legend
			| title=Charles Darwin: vol. 1 Voyaging
	| URL = http://web.archive.org/web/19980119234102/natcenscied.org/PADREV.HTM
			| location=London
	| Publisher = National Center for Science Education}} Retrieved on ]
			| publisher=Jonathan Cape
	*{{Harvard reference
			| isbn=978-1-84413-314-7
	| Surname1 = Paul
	| Given1 = Diane B.
	| Chapter = Darwin, social Darwinism and eugenics
	| Editor = Hodge, Jonathan and Radick, Gregory
	| Year = 2003
	| Title = The Cambridge Companion to Darwin
	| Publisher = Cambridge University Press
	| Location = Cambridge
	| Pages = 214–239
	| ID = ISBN 0-521-77730-5
	}}		}}
			* {{cite book
	*{{Harvard reference
			| last=Browne
	| Surname1 = Reilly
			| first=E. Janet
	| Given1 = Philip
	| Year = 1991		| year=2002
			| title=Charles Darwin: vol. 2 The Power of Place
	| Title = The surgical solution: a history of involuntary sterilization in the United States
			| location=London
	| Publisher = Johns Hopkins University Press
			| publisher=Jonathan Cape
	| Location = Baltimore }}
			| isbn=978-0-7126-6837-8
	*{{Harvard reference
			}}
	| Surname1 = Rothman
			* {{cite book
	| Given1 = Robert
			| last=Darwin
	| Year = 2000 <!--see http://web.archive.org/web/*/http://www.rit.edu/~rhrsbi/GalapagosPages/DarwinFinch.html -->
			| first=Charles
	| Title = Darwin's finches
			| editor-last=Henslow
	| URL = http://www.rit.edu/~rhrsbi/GalapagosPages/DarwinFinch.html }} Retrieved on ]
			| editor-first=J S
	*{{Harvard reference
			| date=1 December 1835
	| Surname1 = Scott
			| title=
	| Given1 = Michon
			| location=Cambridge
	| Year = 2006
			| publisher=
	| Title = Joseph Hooker
			| url=http://darwin-online.org.uk/content/frameset?pageseq=1&itemID=F1&viewtype=side
	| URL = http://www.strangescience.net/jhooker.htm }} Retrieved on ]
			| access-date=27 January 2022
	*{{Harvard reference
			| archive-date=4 October 2022
	| Surname1 = Smith
			| archive-url=https://web.archive.org/web/20221004015917/http://darwin-online.org.uk/content/frameset?pageseq=1&itemID=F1&viewtype=side
	| Given1 = Charles H.
			| url-status=live
	| Year = 1999
			}}
	| Title = Alfred Russel Wallace on Spiritualism, Man, and Evolution: An Analytical Essay
			* {{cite book
	| URL = http://www.wku.edu/~smithch/essays/ARWPAMPH.htm }} Retrieved on ]
			| last=Darwin
	*{{Harvard reference
			| first=Charles
	| Surname1 = Sulloway
			| year=1837
	| Given1 = Frank J.
			| title=Notebook B: (Transmutation of species)
	| Chapter = The ''Beagle'' collections of Darwin's finches (Geospizinae)
			| publisher=Darwin Online
	| Year = 1982
			| id=CUL-DAR121
	| Title = Bulletin of the British Museum (Natural History) Historical Series 43, No. 2: pp 49–94
			| url=http://darwin-online.org.uk/content/frameset?viewtype=side&itemID=CUL-DAR121.-&pageseq=1
	| Publisher = The British Museum
			| access-date=20 December 2008
	| Location = London
			| archive-date=8 February 2009
	| URL = http://darwin-online.org.uk/content/frameset?itemID=A86&viewtype=image&pageseq=1 }} Retrieved on ]
			| archive-url=https://web.archive.org/web/20090208160457/http://darwin-online.org.uk/content/frameset?itemID=CUL-DAR121.-&viewtype=side&pageseq=1
	*{{Harvard reference
			| url-status=live
	| Surname1 = Sweet
			}}
	| Given1 = William
			* {{cite book
	| Year = 2004
			| last=Darwin
	| Title = Herbert Spencer
			| first=Charles
	| Publisher = Internet Encyclopedia of Philosophy
			| year=1839
	| URL = http://www.iep.utm.edu/s/spencer.htm }} Retrieved on ]
			| title=Narrative of the surveying voyages of His Majesty's Ships Adventure and Beagle between the years 1826 and 1836, describing their examination of the southern shores of South America, and the Beagle's circumnavigation of the globe. Journal and remarks. 1832–1836.
	*{{Harvard reference
			| location=London
	| Surname1 = Wilkins
			| publisher=Henry Colburn
	| Given1 = John S.
			| volume=III
	| Year = 1997
			| url=http://darwin-online.org.uk/content/frameset?itemID=F10.3&viewtype=text&pageseq=1
	| Title = Evolution and Philosophy: Does evolution make might right?
			| access-date=24 October 2008
	| URL = http://www.talkorigins.org/faqs/evolphil/social.html
			| archive-date=15 August 2012
	| Publisher = TalkOrigins}} Retrieved on ]
			| archive-url=https://web.archive.org/web/20120815220204/http://darwin-online.org.uk/content/frameset?itemID=F10.3&viewtype=text&pageseq=1
	*{{Harvard reference
			| url-status=live
	| Surname1 = van Wyhe
			}}
	| Given1 = John
			* {{cite book
	| Year = 2006
			| last=Darwin
	| Title = Charles Darwin: gentleman naturalist: A biographical sketch
			| first=Charles
	| URL = http://darwin-online.org.uk/darwin.html }} Retrieved on ]
			| year=1842
	*{{Harvard reference
			| publication-date=1909
	| Surname1 = von Sydow
			| contribution=Pencil Sketch of 1842
	| Given1 = Momme
			| contribution-url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F1556&pageseq=33
	| Year = 2005
			| editor-last=Darwin
	| Chapter = (pp. 141-156)
			| editor-first=Francis
	| URL =
			| editor-link=Francis Darwin
	| Editor = David M. Knight, Matthew D. Eddy
			| title=The foundations of The origin of species: Two essays written in 1842 and 1844.
	| Title = Science and Beliefs: From Natural Philosophy to Natural Science, 1700–1900
			| publisher=Cambridge University Press
	| Publisher = Ashgate
			| url=http://darwin-online.org.uk/content/frameset?itemID=F1556&viewtype=text&pageseq=1
	| Location = Burlington
			| isbn=978-0-548-79998-7
	| Pages = 141–156
			| access-date=13 December 2006
	| ID = ISBN 0-7546-3996-7
			| archive-date=29 September 2011
	}} Retrieved on ]
			| archive-url=https://web.archive.org/web/20110929181520/http://darwin-online.org.uk/content/frameset?itemID=F1556&viewtype=text&pageseq=1
	*{{Harvard reference
			| url-status=live
	| Surname1 = Yates
			}}
	| Given1 = Simon
			* {{cite book
	| Year = 2003
			| last=Darwin
	| Title = The Lady Hope Story: A Widespread Falsehood
			| first=Charles
	| URL = http://www.talkorigins.org/faqs/hope.html
			| year=1845
	| Publisher = TalkOrigins }} Retrieved on ]
			| title=Journal of researches into the natural history and geology of the countries visited during the voyage of H.M.S. Beagle round the world, under the Command of Capt. Fitz Roy, R.N. 2d edition
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/content/frameset?itemID=F20&viewtype=text&pageseq=1
			| access-date=24 October 2008
			| archive-date=17 September 2011
			| archive-url=https://web.archive.org/web/20110917084240/http://darwin-online.org.uk/content/frameset?itemID=F20&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1859
			| title=On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life
			| edition=1st
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=1
			| access-date=24 October 2008
			| isbn=978-1-4353-9386-8
			| archive-date=5 October 2008
			| archive-url=https://web.archive.org/web/20081005185317/http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1871
			| title=The Descent of Man, and Selection in Relation to Sex
			| edition=1st
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/EditorialIntroductions/Freeman_TheDescentofMan.html
			| access-date=24 October 2008
			| isbn=978-0-8014-2085-6
			| archive-date=12 July 2011
			| archive-url=https://web.archive.org/web/20110712194932/http://darwin-online.org.uk/EditorialIntroductions/Freeman_TheDescentofMan.html
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1872
			| title=The Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life
			| edition=6th
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/content/frameset?itemID=F391&viewtype=text&pageseq=1
			| access-date=1 November 2009
			| isbn=978-1-4353-9386-8
			| archive-date=7 January 2010
			| archive-url=https://web.archive.org/web/20100107064907/http://darwin-online.org.uk/content/frameset?itemID=F391&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1874
			| title=The Descent of Man, and Selection in Relation to Sex
			| edition=2nd
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/EditorialIntroductions/Freeman_TheDescentofMan.html
			| access-date=16 January 2016
			| isbn=978-0-8014-2085-6
			| archive-date=12 July 2011
			| archive-url=https://web.archive.org/web/20110712194932/http://darwin-online.org.uk/EditorialIntroductions/Freeman_TheDescentofMan.html
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1887
			| editor-last=Darwin
			| editor-first=Francis
			| editor-link=Francis Darwin
			| title=The life and letters of Charles Darwin, including an autobiographical chapter
			| location=London
			| publisher=John Murray
			| url=http://darwin-online.org.uk/EditorialIntroductions/Freeman_LifeandLettersandAutobiography.html
			| access-date=4 November 2008
			| isbn=978-0-404-08417-2
			| archive-date=5 March 2011
			| archive-url=https://web.archive.org/web/20110305110738/http://darwin-online.org.uk/EditorialIntroductions/Freeman_LifeandLettersandAutobiography.html
			| url-status=live
			}}
			* {{cite book
			| last=Darwin
			| first=Charles
			| year=1958
			| editor-last=Barlow
			| editor-first=Nora
			| editor-link=Nora Barlow
			| title=The Autobiography of Charles Darwin 1809–1882. With the original omissions restored. Edited and with appendix and notes by his granddaughter Nora Barlow
			| url=http://darwin-online.org.uk/content/frameset?pageseq=1&itemID=F1497&viewtype=text
			| location=London
			| publisher=Collins
			| access-date=28 September 2013
			| archive-date=16 August 2013
			| archive-url=https://web.archive.org/web/20130816093152/http://darwin-online.org.uk/content/frameset?itemID=F1497&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{Cite book
			| last=Darwin
			| first=Charles
			| year=2006
			| contribution=Journal
			| contribution-url=http://darwin-online.org.uk/content/frameset?viewtype=side&itemID=CUL-DAR158.1-76&pageseq=1
			| editor-last=van Wyhe
			| editor-first=John
			| title=Darwin's personal 'Journal' (1809–1881)
			| publisher=Darwin Online
			| id=CUL-DAR158.1–76
			| url=http://darwin-online.org.uk/EditorialIntroductions/vanWyhe_JournalDAR158.html
			| access-date=20 December 2008
			| archive-date=24 December 2008
			| archive-url=https://web.archive.org/web/20081224083758/http://darwin-online.org.uk/EditorialIntroductions/vanWyhe_JournalDAR158.html
			| url-status=live
			}}
			* {{cite book
			| last1=Darwin
			| first1=Charles
			| last2=Costa
			| first2=James T.
			| year=2009
			| title=The Annotated Origin: A Facsimile of the First Edition of On the Origin of Species Annotated by James T. Costa
			| isbn=978-0-674-03281-1
			| location=Cambridge, Massachusetts, and London, England
			| publisher=Belknap Press of Harvard University Press
			| url=https://archive.org/details/annotatedoriginf00darw
			}}
			* {{cite book
			| last1=Desmond
			| first1=Adrian
			| author-link=Adrian Desmond
			| last2=Moore
			| first2=James
			| author2-link=James Moore (biographer)
			| year=1991
			| title=Darwin
			| location=London
			| publisher=Michael Joseph, Penguin Group
			| isbn=978-0-7181-3430-3}}
			* {{cite ODNB
			| last=Desmond
			| first=Adrian
			| last2=Moore
			| first2=James
			| last3=Browne
			| first3=Janet
			| year=2004
			| title=Darwin, Charles Robert
			| location=Oxford, England
			| doi=10.1093/ref:odnb/7176
			| isbn=978-0-19-861411-1
			}}
			* {{cite book
			| last1=Desmond
			| first1=Adrian
			| last2=Moore
			| first2=James
			| last3=Browne
			| first3=Janet
			| year=2007
			| title=Charles Darwin (Very Interesting People)
			| publisher=Oxford University Press
			| location=Oxford, England
			| isbn=978-0-19-921354-2
			}}
			* {{cite book
			| last1=Desmond
			| first1=Adrian
			| last2=Moore
			| first2=James
			| title=Darwin's sacred cause : race, slavery and the quest for human origins
			| publisher=Allen Lane
			| location=London
			| year=2009
			| isbn=978-1-84614-035-8
			| url-access=registration
			| url=https://archive.org/details/darwinssacredcau0000desm
			}}
			* {{cite journal
			| last=Eldredge
			| first=Niles
			| author-link=Niles Eldredge
			| year=2006
			| title=Confessions of a Darwinist
			| periodical=The Virginia Quarterly Review
			| issue=Spring 2006
			| pages=32–53
			| url=http://www.vqronline.org/articles/2006/spring/eldredge-confessions-darwinist/
			| access-date=4 November 2008
			| archive-date=24 December 2013
			| archive-url=https://web.archive.org/web/20131224110620/http://www.vqronline.org/articles/2006/spring/eldredge-confessions-darwinist/
			| url-status=dead
			}}
			* {{cite book
			| last=FitzRoy
			| first=Robert
			| author-link=Robert Fitzroy
			| year=1839
			| title=Voyages of the Adventure and Beagle, Volume II
			| location=London
			| publisher=Henry Colburn
			| url=http://darwin-online.org.uk/content/frameset?itemID=F10.2&viewtype=text&pageseq=1
			| access-date=4 November 2008
			| archive-date=5 May 2011
			| archive-url=https://web.archive.org/web/20110505173517/http://darwin-online.org.uk/content/frameset?itemID=F10.2&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite book
			| last=Freeman
			| first=R. B.
			| author-link=R. B. Freeman
			| year=1977
			| title=The Works of Charles Darwin: An Annotated Bibliographical Handlist
			| location=Folkestone
			| publisher=Wm Dawson & Sons Ltd
			| url=http://darwin-online.org.uk/content/frameset?pageseq=4&itemID=A1&viewtype=side
			| access-date=4 November 2008
			| isbn=978-0-208-01658-4
			| url-access=
			| archive-date=4 October 2022
			| archive-url=https://web.archive.org/web/20221004023516/http://darwin-online.org.uk/content/frameset?pageseq=4&itemID=A1&viewtype=side
			| url-status=live
			}}
			* {{cite book
			| last=Freeman
			| first=R. B.
			| title=Charles Darwin: A companion
			| publisher=]
			| year=2007
			| edition=2nd online
			| url=http://darwin-online.org.uk/content/frameset?itemID=A27b&viewtype=text&pageseq=114
			| pages=107, 109
			| access-date=25 December 2014
			| archive-date=25 December 2014
			| archive-url=https://web.archive.org/web/20141225163344/http://darwin-online.org.uk/content/frameset?itemID=A27b&viewtype=text&pageseq=114
			| url-status=live
			}}
			* {{cite journal
			| last=Herbert
			| first=Sandra
			| year=1980
			| title=The red notebook of Charles Darwin
			| journal=Bulletin of the British Museum (Natural History), Historical Series
			| volume=7
			| issue=7 (24 April)
			| pages=1–164
			| doi=10.5962/p.272299
			| url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F1583e&pageseq=1
			| access-date=11 January 2009
			| archive-date=11 July 2007
			| archive-url=https://web.archive.org/web/20070711050113/http://darwin-online.org.uk/content/frameset?itemID=F1583e&viewtype=text&pageseq=1
			| url-status=live
			| doi-access=free
			}}
			* {{cite journal
			| last=Herbert
			| first=Sandra
			| year=1991
			| title=Charles Darwin as a prospective geological author
			| journal=British Journal for the History of Science
			| issue=2
			| pages=159–192
			| url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=A342&pageseq=1
			| access-date=24 October 2008
			| doi=10.1017/S0007087400027060
			| volume=24
			| s2cid=143748414
			| archive-date=29 March 2017
			| archive-url=https://web.archive.org/web/20170329133528/http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=A342&pageseq=1
			| url-status=live
			}}
			*{{cite book|last=Holliday |first=Trenton |title=Cro-Magnon: The Story of the Last Ice Age People of Europe |publisher= Columbia University Press|location =NewYork |year=2021|isbn=978-0-231-20497-2}}
			* {{cite book
			| last1=Huxley
			| first1=Julian
			| author-link=Julian Huxley
			| last2=Kettlewell
			| first2=H.B.D.
			| author-link2=Bernard Kettlewell
			| title=Charles Darwin and His World
			| url=https://archive.org/details/charlesdarwinhis0000huxl_y9d3
			| url-access=registration
			| publisher=the Viking Press
			| location=New York
			| year=1965
			}}
			* {{cite book
			| last=Keynes
			| first=Richard
			| author-link=Richard Keynes
			| year=2000
			| title=Charles Darwin's zoology notes & specimen lists from H.M.S. Beagle
			| publisher=Cambridge University Press
			| url=http://darwin-online.org.uk/content/frameset?itemID=F1840&viewtype=text&pageseq=1
			| access-date=22 November 2008
			| isbn=978-0-521-46569-4
			| archive-date=5 December 2008
			| archive-url=https://web.archive.org/web/20081205002654/http://darwin-online.org.uk/content/frameset?itemID=F1840&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite book
			| last=Keynes
			| first=Richard
			| year=2001
			| title=Charles Darwin's Beagle Diary
			| publisher=Cambridge University Press
			| url=http://darwin-online.org.uk/content/frameset?itemID=F1925&viewtype=text&pageseq=1
			| access-date=24 October 2008
			| isbn=978-0-521-23503-7
			| archive-date=4 June 2012
			| archive-url=https://archive.today/20120604052049/http://darwin-online.org.uk/content/frameset?itemID=F1925&viewtype=text&pageseq=1
			| url-status=live
			}}
			* {{cite web
			| last=Kotzin
			| first=Daniel
			| year=2004
			| title=Point-Counterpoint: Social Darwinism
			| publisher=Columbia American History Online
			| url=http://caho-test.cc.columbia.edu/pcp/14008.html
			| access-date=22 November 2008
			| url-status=dead
			| archive-url=https://web.archive.org/web/20110719072856/http://caho-test.cc.columbia.edu/pcp/14008.html
			| archive-date=19 July 2011
			}}
			* {{cite book| last=Larson| first=Edward J.| author-link=Edward Larson| title=Evolution: The Remarkable History of a Scientific Theory| publisher=Modern Library| year=2004| isbn=978-0-679-64288-6| url=https://archive.org/details/evolutionremarka00lars}}
			* {{cite web
			| last=Leff
			| first=David
			| year=2000
			| title=AboutDarwin.com
			| url=http://www.aboutdarwin.com/index.html
			| edition=2000–2008
			| access-date=30 December 2008
			| archive-url=https://web.archive.org/web/20130828111301/http://www.aboutdarwin.com/index.html
			| archive-date=28 August 2013
			| url-status=dead
			| df=dmy-all
			}}
			* {{cite journal
			| last=Leifchild
			| date=19 November 1859
			| title=Review of 'Origin'
			| periodical=]
			| issue=1673
			| url=http://darwin-online.org.uk/content/frameset?viewtype=image&itemID=CUL-DAR226.1.8&pageseq=1
			| access-date=22 November 2008
			| archive-date=5 December 2008
			| archive-url=https://web.archive.org/web/20081205002714/http://darwin-online.org.uk/content/frameset?viewtype=image&itemID=CUL-DAR226.1.8&pageseq=1
			| url-status=live
			}}
			* {{cite journal
			| last=Miles
			| first=Sara Joan
			| year=2001
			| title=Charles Darwin and Asa Gray Discuss Teleology and Design
			| journal=]
			| volume=53
			| pages=196–201
			| url=http://www.asa3.org/ASA/PSCF/2001/PSCF9-01Miles.html
			| access-date=22 November 2008
			| archive-date=5 April 2020
			| archive-url=https://web.archive.org/web/20200405172817/http://www.asa3.org/ASA/PSCF/2001/PSCF9-01Miles.html
			| url-status=dead
			}}
			* {{cite news
			| last=Moore
			| first=James
			| author-link=James Moore (biographer)
			| year=2005
			| title=Darwin&nbsp;– A 'Devil's Chaplain'?
			| publisher=American Public Media
			| url=http://speakingoffaith.publicradio.org/programs/darwin/moore-devilschaplain.pdf
			| access-date=22 November 2008
			| url-status=dead
			| archive-url=https://web.archive.org/web/20080227014518/http://speakingoffaith.publicradio.org/programs/darwin/moore-devilschaplain.pdf
			| archive-date=27 February 2008
			}}
			* {{cite news
			| last=Moore
			| first=James
			| year=2006
			| title=Evolution and Wonder&nbsp;– Understanding Charles Darwin
			| series=Speaking of Faith (Radio Program)
			| publisher=American Public Media
			| url=http://speakingoffaith.publicradio.org/programs/darwin/transcript.shtml
			| access-date=22 November 2008
			| url-status=dead
			| archive-url=https://web.archive.org/web/20081222020720/http://speakingoffaith.publicradio.org/programs/darwin/transcript.shtml
			| archive-date=22 December 2008
			}}
			* {{cite book
			| last=Owen
			| first=Richard
			| author-link=Richard Owen
			| year=1840
			| editor-last=Darwin
			| editor-first=C. R.
			| title=Fossil Mammalia Part 1
			| series=The zoology of the voyage of H.M.S. Beagle
			| location=London
			| publisher=Smith Elder and Co}}
			* {{cite book
			| last=Paul
			| first=Diane B.
			| year=2003
			| contribution=Darwin, social Darwinism and eugenics
			| editor-last=Hodge
			| editor-first=Jonathan
			| editor2-last=Radick
			| editor2-first=Gregory
			| title=The Cambridge Companion to Darwin
			| url=https://archive.org/details/cambridgecompani00hodg_248
			| url-access=limited
			| publisher=Cambridge University Press
			| pages=–239
			| isbn=978-0-521-77730-8
			}}
			* {{cite book
			| last=Radick
			| first=Gregory
			| chapter=Darwin and Humans
			| editor-last=Ruse
			| editor-first=Michael
			| title=The Cambridge Encyclopedia of Darwin and Evolutionary Thought
			| publisher=Cambridge University Press
			| year=2013
			| pages=173–181
			}}
			* {{cite web
			| last=Smith
			| first=Charles H.
			| title=Alfred Russel Wallace on Spiritualism, Man, and Evolution: An Analytical Essay
			| year=1999
			| url=http://www.wku.edu/~smithch/essays/ARWPAMPH.htm
			| access-date=7 December 2008
			| archive-date=5 December 2008
			| archive-url=https://web.archive.org/web/20081205020823/http://www.wku.edu/~smithch/essays/ARWPAMPH.htm
			| url-status=live
			}}
			* {{cite journal
			| last=Sulloway
			| first=Frank J.
			| author-link=Frank Sulloway
			| year=1982
			| title=Darwin and His Finches: The Evolution of a Legend
			| journal=Journal of the History of Biology
			| volume=15
			| issue=1
			| pages=1–53
			| url=http://www.sulloway.org/Finches.pdf
			| access-date=9 December 2008
			| doi=10.1007/BF00132004
			| citeseerx=10.1.1.458.3975
			| s2cid=17161535
			| archive-date=16 December 2008
			| archive-url=https://web.archive.org/web/20081216211931/http://www.sulloway.org/Finches.pdf
			| url-status=dead
			}}
			* {{cite web
			| last=Sweet
			| first=William
			| title=Herbert Spencer
			| publisher=Internet Encyclopedia of Philosophy
			| year=2004
			| url=http://www.iep.utm.edu/spencer/
			| access-date=16 December 2008
			| archive-date=28 May 2010
			| archive-url=https://web.archive.org/web/20100528161329/http://www.iep.utm.edu/spencer/
			| url-status=live
			}}
			* {{cite web
			| last=Wilkins
			| first=John S.
			| year=1997
			| title=Evolution and Philosophy: Does evolution make might right?
			| publisher=]
			| url=http://www.talkorigins.org/faqs/evolphil/social.html
			| access-date=22 November 2008
			| archive-date=14 May 2011
			| archive-url=https://web.archive.org/web/20110514095809/http://www.talkorigins.org/faqs/evolphil/social.html
			| url-status=live
			}}
			* {{cite book
			| last=Wilkins
			| first=John S.
			| year=2008
			| contribution=Darwin
			| editor-last=Tucker
			| editor-first=Aviezer
			| title=A Companion to the Philosophy of History and Historiography
			| series=Blackwell Companions to Philosophy
			| pages=405–415
			| location=Chichester
			| publisher=Wiley-Blackwell
			| isbn=978-1-4051-4908-2}}
			* {{cite journal
			| last=van Wyhe
			| first=John
			| title=Mind the gap: Did Darwin avoid publishing his theory for many years?
			| journal=Notes and Records of the Royal Society
			| volume=61
			| issue=2
			| pages=177–205
			| date=27 March 2007
			| doi=10.1098/rsnr.2006.0171
			| s2cid=202574857
			| url=http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=A544&pageseq=1
			| access-date=7 February 2008
			| archive-date=11 January 2011
			| archive-url=https://web.archive.org/web/20110111012141/http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=A544&pageseq=1
			| url-status=live
			}}
			* {{cite web
			| last=van Wyhe
			| first=John
			| year=2008
			| title=Charles Darwin: gentleman naturalist: A biographical sketch
			| publisher=Darwin Online
			| url=http://darwin-online.org.uk/darwin.html
			| access-date=17 November 2008
			| archive-date=13 January 2020
			| archive-url=https://web.archive.org/web/20200113061826/http://darwin-online.org.uk/darwin.html
			| url-status=live
			}}
			* {{cite book
			| last=van Wyhe
			| first=John
			| publication-date=1 September 2008
			| year=2008b
			| title=Darwin: The Story of the Man and His Theories of Evolution
			| location=London
			| publisher=Andre Deutsch Ltd
			| isbn=978-0-233-00251-4}}
			* {{cite book
			| last=von Sydow
			| first=Momme
			| year=2005
			| contribution=Darwin&nbsp;– A Christian Undermining Christianity? On Self-Undermining Dynamics of Ideas Between Belief and Science
			| contribution-url=http://www.psych.uni-goettingen.de/abt/1/sydow/von_Sydow_(2005)_Darwin_A_Christian_Undermining_Christianity.pdf
			| editor-last=Knight
			| editor-first=David M.
			| editor2-last=Eddy
			| editor2-first=Matthew D.
			| title=Science and Beliefs: From Natural Philosophy to Natural Science, 1700–1900
			| location=Burlington
			| publisher=Ashgate
			| pages=141–156
			| isbn=978-0-7546-3996-1
			| access-date=16 December 2008
			| archive-url=https://web.archive.org/web/20090326070105/http://www.psych.uni-goettingen.de/abt/1/sydow/von_Sydow_(2005)_Darwin_A_Christian_Undermining_Christianity.pdf
			| archive-date=26 March 2009
			}}
			* {{cite web
			| last=Yates
			| first=Simon
			| year=2003
			| title=The Lady Hope Story: A Widespread Falsehood
			| publisher=]
			| url=http://www.talkorigins.org/faqs/hope.html
			| access-date=15 December 2006
			| archive-date=12 October 2009
			| archive-url=https://web.archive.org/web/20091012194435/http://www.talkorigins.org/faqs/hope.html
			| url-status=live
			}}
			{{refend}}
	==External links==		==External links==
			{{Library resources box
	{{sisterlinks|Charles Darwin}}
			| onlinebooks = yes
	*
			| by = yes
	*
			| viaf = 27063124
	*
			| label = Charles Darwin
	*
			}}
	*
			* {{cite web |title=The Complete Work of Charles Darwin Online |url=https://darwin-online.org.uk/ |access-date=4 March 2024}}
	*
			* {{StandardEbooks|Standard Ebooks URL=https://standardebooks.org/ebooks/charles-darwin}}
	*
			* {{Gutenberg author |id=485| name=Charles Darwin}}
	*
			* {{Internet Archive author |name=Charles Robert Darwin}}
	*
			* {{Librivox author |id=166}}
	*
			* ]&nbsp;– ; Darwin's publications, private papers and bibliography, supplementary works including biographies, obituaries and reviews
	*
			* Full text and notes for complete correspondence to 1867, with summaries of all the rest, and pages of commentary
	*
	*		*
			* {{UK National Archives ID}}
	*
			* View books owned and annotated by at the online Biodiversity Heritage Library.
	*{{dmoz|Science/Biology/History/People/Darwin,_Charles/}}
			* {{BHL author}}
	*
			* in ]
	* , an organisation to celebrate Darwin’s bicentenary in February 2009.
			* {{NPG name}}
			* {{PM20|FID=pe/003703}}
			* – Occasional Papers from RHS Lindley Library, volume 3 July 2010
			* ], 29 April 1882, pp.&nbsp;256,
			* {{IEP|/darwin/}}
	{{Darwin}}		{{Darwin}}
			{{Navboxes
			|title=Links to related articles
	{{evolution}}
			|list1=
			{{Evolution}}
	<!-- Metadata: see ] -->
			{{Ethology}}
			{{Zoology}}
	{{Persondata
			{{Evolutionary psychologists}}
	|NAME= Darwin, Charles Robert
			{{Copley Medallists 1851–1900}}
	|ALTERNATIVE NAMES=
			{{FRS 1839}}
	|SHORT DESCRIPTION= ]
			{{Nonverbal communication}}
	|DATE OF BIRTH= ] ]
	|PLACE OF BIRTH= ], ], ], ]
	|DATE OF DEATH= ] ]
	|PLACE OF DEATH= ], ], ]
	}}		}}
			{{Subject bar
			| commons = y
			| q = y
			| s = y
			| s-search = Author:Charles Robert Darwin
			| wikt = Darwinian
			| d = y
			| d-search = Q160627
			| species_author=yes
			| b = General Biology/Gallery of Biologists/Charles Darwin
			}}
			{{Authority control|suppress=P434}}
	{{DEFAULTSORT:Darwin, Charles}}		{{DEFAULTSORT:Darwin, Charles}}
	]		]
	]		]
	]		]
	]		]
	]		]
			]
			]
			]
			]
			]
			]
			]
			]
			]
	]		]
	]		]
			]
			]
			]
			]
			]
			]
			]
			]
	]		]
			]
	]
			]
			]
			]
			]
	]		]
	]		]
	]		]
	]		]
	]		]
	]		]
	]		]
	]		]
			]
	]
	]		]
	]		]
	]		]
	]		]
			]
			]
	{{Link FA|el}}
			]
	{{Link FA|sl}}
			]
	{{Link FA|sr}}
	{{Link FA|bs}}
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]
	]

---

Latest revision as of 20:17, 7 January 2025

English naturalist and biologist (1809–1882) For other people named Charles Darwin, see Charles Darwin (disambiguation).

Charles DarwinJP FRS FRGS FLS FZS
Darwin, c. 1854, when he was preparing On the Origin of Species
Born	Charles Robert Darwin (1809-02-12)12 February 1809 Shrewsbury, Shropshire, England
Died	19 April 1882(1882-04-19) (aged 73) Down House, Down, Kent, England
Resting place	Westminster Abbey
Education	University of Edinburgh Christ's College, Cambridge (BA, MA)
Known for	Natural selection
Spouse	Emma Wedgwood ​(m. 1839)​
Children	10, including William, Henrietta, George, Francis, Leonard and Horace
Parents	Robert Darwin Susannah Wedgwood
Family	Darwin–Wedgwood
Awards	FRS (1839) Royal Medal (1853) Wollaston Medal (1859) Copley Medal (1864) Pour le Mérite (1867) Baly Medal (1879)
Writing career
Notable works	The Voyage of the Beagle On the Origin of Species The Descent of Man
Scientific career
Fields	Natural history Geology
Institutions	Geological Society of London
Academic advisors	John Stevens Henslow Adam Sedgwick
Author abbrev. (botany)	Darwin
Author abbrev. (zoology)	Darwin
Signature

Charles Robert Darwin (/ˈdɑːrwɪn/ DAR-win; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended from a common ancestor is now generally accepted and considered a fundamental scientific concept. In a joint presentation with Alfred Russel Wallace, he introduced his scientific theory that this branching pattern of evolution resulted from a process he called natural selection, in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding. Darwin has been described as one of the most influential figures in human history and was honoured by burial in Westminster Abbey.

Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh; instead, he helped to investigate marine invertebrates. His studies at the University of Cambridge's Christ's College from 1828 to 1831 encouraged his passion for natural science. However, it was his five-year voyage on HMS Beagle from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported Charles Lyell's concept of gradual geological change. Publication of his journal of the voyage made Darwin famous as a popular author.

Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting the immediate joint submission of both their theories to the Linnean Society of London. Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil.

Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species. By the 1870s, the scientific community and a majority of the educated public had accepted evolution as a fact. However, many initially favoured competing explanations that gave only a minor role to natural selection, and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution. Darwin's scientific discovery is the unifying theory of the life sciences, explaining the diversity of life.

Biography

Early life and education

Further information: Charles Darwin's education and Darwin–Wedgwood family

Darwin was born in Shrewsbury, Shropshire, on 12 February 1809, at his family's home, The Mount. He was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists. Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794), a poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded.

Both families were largely Unitarian, though the Wedgwoods were adopting Anglicanism. Robert Darwin, a freethinker, had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury, but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had a taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending the nearby Anglican Shrewsbury School as a boarder.

Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone, a freed black slave who had accompanied Charles Waterton in the South American rainforest.

In Darwin's second year at the university, he joined the Plinian Society, a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant's investigations of the anatomy and life cycle of marine invertebrates in the Firth of Forth, and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech. One day, Grant praised Lamarck's evolutionary ideas. Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin was rather bored by Robert Jameson's natural-history course, which covered geology – including the debate between neptunism and plutonism. He learned the classification of plants and assisted with work on the collections of the University Museum, one of the largest museums in Europe at the time.

Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge, in January 1828, to study for a Bachelor of Arts degree as the first step towards becoming an Anglican country parson. Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying.

During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens' Illustrations of British entomology (1829–1932).

Through Fox, Darwin became a close friend and follower of botany professor John Stevens Henslow. He met other leading parson-naturalists who saw scientific work as religious natural theology, becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for the ordinary degree.

Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature, explaining adaptation as God acting through laws of nature. He read John Herschel's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described the highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick's geology course, then on 4 August travelled with him to spend a fortnight mapping strata in Wales.

Survey voyage on HMS Beagle

Further information: Second voyage of HMS Beagle

After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth. He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS Beagle with captain Robert FitzRoy, a position for a gentleman rather than "a mere collector". The ship was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II, to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution.

After delays, the voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations. At intervals during the voyage, his specimens were sent to Cambridge together with letters including a copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during a calm spell.

On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell's Principles of Geology, which set out uniformitarian concepts of land slowly rising or falling over immense periods, and Darwin saw things Lyell's way, theorising and thinking of writing a book on geology. When they reached Brazil, Darwin was delighted by the tropical forest, but detested the sight of slavery there, and disputed this issue with FitzRoy.

The survey continued to the south in Patagonia. They stopped at Bahía Blanca, and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like a giant version of the armour on local armadillos. From a jaw and tooth he identified the gigantic Megatherium, then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest. In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles.

On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at a time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species.

Three Fuegians on board, who had been seized during the first Beagle voyage then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England.

Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel-beds stranded above high tide. High in the Andes he saw seashells and several fossil trees that had grown on a sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls.

On the geologically new Galápagos Islands, Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia, the marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aborigines "good-humoured & pleasant", their numbers depleted by European settlement.

FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising. FitzRoy began writing the official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history.

In Cape Town, South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries, the replacement of extinct species by others" as "a natural in contradistinction to a miraculous process". When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on the origin of species".

Without telling Darwin, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society, and reported in magazines, including The Athenaeum. Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have a great name among the Naturalists of Europe".

Inception of Darwin's evolutionary theory

Further information: Inception of Darwin's theory

On 2 October 1836, Beagle anchored at Falmouth, Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to Cambridge to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be a self-funded gentleman scientist, and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage.

Charles Lyell eagerly met Darwin for the first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen, who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium Darwin had identified, a near complete skeleton of the unknown Scelidotherium and a hippopotamus-sized rodent-like skull named Toxodon resembling a giant capybara. The armour fragments were actually from Glyptodon, a huge armadillo-like creature, as Darwin had initially thought. These extinct creatures were related to living species in South America.

In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into the Narrative were resolved at the end of the month when FitzRoy accepted Broderip's advice to make it a separate volume, and Darwin began work on his Journal and Remarks.

Darwin's first paper showed that the South American landmass was slowly rising. With Lyell's enthusiastic backing, he read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to the Zoological Society. The ornithologist John Gould soon announced that the Galápagos birds that Darwin had thought a mixture of blackbirds, "gros-beaks" and finches, were, in fact, twelve separate species of finches. On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas.

Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage, who described God as a programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and a close friend of the writer Harriet Martineau, who promoted the Malthusianism that underpinned the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the radical implications of transmutation of species, promoted by Grant and younger surgeons influenced by Geoffroy. Transmutation was anathema to Anglicans defending social order, but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species.

Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought was a "wren" was in the finch group. Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands. The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards.

By mid-March 1837, barely six months after his return to England, Darwin was speculating in his Red Notebook on the possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal Macrauchenia, which resembled a giant guanaco, a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations – explaining the variations he had observed in Galápagos tortoises, mockingbirds, and rheas. He sketched branching descent, and then a genealogical branching of a single evolutionary tree, in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent lineages progressing to higher forms.

Overwork, illness, and marriage

Further information: Health of Charles Darwin

While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his Journal, he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained a Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle, a sum equivalent to about £115,000 in 2021. He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher. As the Victorian era began, Darwin pressed on with writing his Journal, and in August 1837 began correcting printer's proofs.

As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable palpitation of the heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at Maer Hall, Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood, nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been the work of earthworms, inspiring "a new & important theory" on their role in soil formation, which Darwin presented at the Geological Society on 1 November 1837. His Journal was printed and ready for publication by the end of February 1838, as was the first volume of the Narrative, but FitzRoy was still working hard to finish his own volume.

William Whewell pushed Darwin to take on the duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in selective breeding such as farmers and pigeon fanciers. Over time, his research drew on information from his relatives and children, the family butler, neighbours, colonists and former shipmates. He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour.

The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils, palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of Darwin's illness remained unknown, and attempts at treatment had only ephemeral success.

On 23 June, he took a break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a proglacial lake.

Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry". Advantages under "Marry" included "constant companion and a friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time". Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation. He married Emma on 29 January 1839 and they were the parents of ten children, seven of whom survived to adulthood.

Malthus and natural selection

Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus's An Essay on the Principle of Population. On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what is known as a Malthusian catastrophe. Darwin was well-prepared to compare this to Augustin de Candolle's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that the "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones." This would result in the formation of new species. As he later wrote in his Autobiography:

In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate the struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work...

By mid-December, Darwin saw a similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected", thinking this comparison "a beautiful part of my theory". He later called his theory natural selection, an analogy with what he termed the "artificial selection" of selective breeding.

On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in the afterlife. While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street, then moved his "museum" in over Christmas. On 24 January 1839, Darwin was elected a Fellow of the Royal Society (FRS).

On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home.

Geology books, barnacles, evolutionary research

Further information: Development of Darwin's theory

Darwin now had the framework of his theory of natural selection "by which to work", as his "prime hobby". His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory. For fifteen years this work was in the background to his main occupation of writing on geology and publishing expert reports on the Beagle collections, in particular, the barnacles.

The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed Mr. Arthrobalanus. His confusion over the relationship of this species (Cryptophialus minutus) to other barnacles caused him to fixate on the systematics of the taxa. He wrote his first examination of the species in 1846 but did not formally describe it until 1854.

FitzRoy's long-delayed Narrative was published in May 1839. Darwin's Journal and Remarks got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing a beginning to each crop of species".

Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection. To escape the pressures of London, the family moved to rural Down House in Kent in September. On 11 January 1844, Darwin mentioned his theorising to the botanist Joseph Dalton Hooker, writing with melodramatic humour "it is like confessing a murder". Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject."

By July, Darwin had expanded his "sketch" into a 230-page "Essay", to be expanded with his research results if he died prematurely. In November, the anonymously published sensational best-seller Vestiges of the Natural History of Creation brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists.

Darwin completed his third geological book in 1846. He now renewed a fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation.

In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully's Malvern spa and was surprised to find some benefit from hydrotherapy. Then, in 1851, his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises.

In eight years of work on barnacles, Darwin's theory helped him to find "homologies" showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites, showing an intermediate stage in evolution of distinct sexes. In 1853, it earned him the Royal Society's Royal Medal, and it made his reputation as a biologist. Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before." In 1854, he became a Fellow of the Linnean Society of London, gaining postal access to its library. He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in the economy of nature".

Publication of the theory of natural selection

Further information: Publication of Darwin's theory

By the start of 1856, Darwin was investigating whether eggs and seeds could survive travel across seawater to spread species across oceans. Hooker increasingly doubted the traditional view that species were fixed, but their young friend Thomas Henry Huxley was still firmly against the transmutation of species. Lyell was intrigued by Darwin's speculations without realising their extent. When he read a paper by Alfred Russel Wallace, "On the Law which has Regulated the Introduction of New Species", he saw similarities with Darwin's thoughts and urged him to publish to establish precedence.

Though Darwin saw no threat, on 14 May 1856 he began writing a short paper. Finding answers to difficult questions held him up repeatedly, and he expanded his plans to a "big book on species" titled Natural Selection, which was to include his "note on Man". He continued his research, obtaining information and specimens from naturalists worldwide, including Wallace who was working in Borneo.

In mid-1857, he added a section heading, "Theory applied to Races of Man", but did not add text on this topic. On 5 September 1857, Darwin sent the American botanist Asa Gray a detailed outline of his ideas, including an abstract of Natural Selection, which omitted human origins and sexual selection. In December, Darwin received a letter from Wallace asking if the book would examine human origins. He responded that he would avoid that subject, "so surrounded with prejudices", while encouraging Wallace's theorising and adding that "I go much further than you."

Darwin's book was only partly written when, on 18 June 1858, he received a paper from Wallace describing natural selection. Shocked that he had been "forestalled", Darwin sent it on that day to Lyell, as requested by Wallace, and although Wallace had not asked for publication, Darwin suggested he would send it to any journal that Wallace chose. His family was in crisis, with children in the village dying of scarlet fever, and he put matters in the hands of his friends. After some discussion, with no reliable way of involving Wallace, Lyell and Hooker decided on a joint presentation at the Linnean Society on 1 July of On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection. On the evening of 28 June, Darwin's baby son died of scarlet fever after almost a week of severe illness, and he was too distraught to attend.

There was little immediate attention to this announcement of the theory; the president of the Linnean Society remarked in May 1859 that the year had not been marked by any revolutionary discoveries. Only one review rankled enough for Darwin to recall it later; Professor Samuel Haughton of Dublin claimed that "all that was new in them was false, and what was true was old". Darwin struggled for thirteen months to produce an abstract of his "big book", suffering from ill health but getting constant encouragement from his scientific friends. Lyell arranged to have it published by John Murray.

On the Origin of Species proved unexpectedly popular, with the entire stock of 1,250 copies oversubscribed when it went on sale to booksellers on 22 November 1859. In the book, Darwin set out "one long argument" of detailed observations, inferences and consideration of anticipated objections. In making the case for common descent, he included evidence of homologies between humans and other mammals. Having outlined sexual selection, he hinted that it could explain differences between human races. He avoided explicit discussion of human origins, but implied the significance of his work with the sentence; "Light will be thrown on the origin of man and his history." His theory is simply stated in the introduction:

As many more individuals of each species are born than can possibly survive; and as, consequently, there is a frequently recurring struggle for existence, it follows that any being, if it vary however slightly in any manner profitable to itself, under the complex and sometimes varying conditions of life, will have a better chance of surviving, and thus be naturally selected. From the strong principle of inheritance, any selected variety will tend to propagate its new and modified form.

At the end of the book, he concluded that:

There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been, and are being, evolved.

The last word was the only variant of "evolved" in the first five editions of the book. "Evolutionism" at that time was associated with other concepts, most commonly with embryological development. Darwin first used the word evolution in The Descent of Man in 1871, before adding it in 1872 to the 6th edition of The Origin of Species.

Responses to publication

Further information: Reactions to On the Origin of Species

The book aroused international interest, with less controversy than had greeted the popular and less scientific Vestiges of the Natural History of Creation. Though Darwin's illness kept him away from the public debates, he eagerly scrutinised the scientific response, commenting on press cuttings, reviews, articles, satires and caricatures, and corresponded on it with colleagues worldwide. The book did not explicitly discuss human origins, but included a number of hints about the animal ancestry of humans from which the inference could be made.

The first review asked, "If a monkey has become a man – what may not a man become?" It said this should be left to theologians as being too dangerous for ordinary readers. Among early favourable responses, Huxley's reviews swiped at Richard Owen, leader of the scientific establishment which Huxley was trying to overthrow.

In April, Owen's review attacked Darwin's friends and condescendingly dismissed his ideas, angering Darwin, but Owen and others began to promote ideas of supernaturally guided evolution. Patrick Matthew drew attention to his 1831 book which had a brief appendix suggesting a concept of natural selection leading to new species, but he had not developed the idea.

The Church of England's response was mixed. Darwin's old Cambridge tutors Sedgwick and Henslow dismissed the ideas, but liberal clergymen interpreted natural selection as an instrument of God's design, with the cleric Charles Kingsley seeing it as "just as noble a conception of Deity". In 1860, the publication of Essays and Reviews by seven liberal Anglican theologians diverted clerical attention from Darwin. Its ideas, including higher criticism, were attacked by church authorities as heresy. In it, Baden Powell argued that miracles broke God's laws, so belief in them was atheistic, and praised "Mr Darwin's masterly volume [supporting] the grand principle of the self-evolving powers of nature".

Asa Gray discussed teleology with Darwin, who imported and distributed Gray's pamphlet on theistic evolution, Natural Selection is not inconsistent with natural theology. The most famous confrontation was at the public 1860 Oxford evolution debate during a meeting of the British Association for the Advancement of Science, where the Bishop of Oxford Samuel Wilberforce, though not opposed to transmutation of species, argued against Darwin's explanation and human descent from apes. Joseph Hooker argued strongly for Darwin, and Thomas Huxley's legendary retort, that he would rather be descended from an ape than a man who misused his gifts, came to symbolise a triumph of science over religion.

Even Darwin's close friends Gray, Hooker, Huxley and Lyell still expressed various reservations but gave strong support, as did many others, particularly younger naturalists. Gray and Lyell sought reconciliation with faith, while Huxley portrayed a polarisation between religion and science. He campaigned pugnaciously against the authority of the clergy in education, aiming to overturn the dominance of clergymen and aristocratic amateurs under Owen in favour of a new generation of professional scientists. Owen's claim that brain anatomy proved humans to be a separate biological order from apes was shown to be false by Huxley in a long-running dispute parodied by Kingsley as the "Great Hippocampus Question", and discredited Owen. In response to objections that the origin of life was unexplained, Darwin pointed to acceptance of Newton's law even though the cause of gravity was unknown. Despite criticisms and reservations related to this topic, he nevertheless proposed a prescient idea in an 1871 letter to Hooker in which he suggested the origin of life may have occurred in a "warm little pond".

Darwinism became a movement covering a wide range of evolutionary ideas. In 1863, Lyell's Geological Evidences of the Antiquity of Man popularised prehistory, though his caution on evolution disappointed Darwin. Weeks later Huxley's Evidence as to Man's Place in Nature showed that anatomically, humans are apes, then The Naturalist on the River Amazons by Henry Walter Bates provided empirical evidence of natural selection. Lobbying brought Darwin Britain's highest scientific honour, the Royal Society's Copley Medal, awarded on 3 November 1864. That day, Huxley held the first meeting of what became the influential "X Club" devoted to "science, pure and free, untrammelled by religious dogmas". By the end of the decade, most scientists agreed that evolution occurred, but only a minority supported Darwin's view that the chief mechanism was natural selection.

The Origin of Species was translated into many languages, becoming a staple scientific text attracting thoughtful attention from all walks of life, including the "working men" who flocked to Huxley's lectures. Darwin's theory resonated with various movements at the time and became a key fixture of popular culture. Cartoonists parodied animal ancestry in an old tradition of showing humans with animal traits, and in Britain, these droll images served to popularise Darwin's theory in an unthreatening way. While ill in 1862, Darwin began growing a beard, and when he reappeared in public in 1866, caricatures of him as an ape helped to identify all forms of evolutionism with Darwinism.

Othniel C. Marsh, America's first palaeontologist, was the first to provide solid fossil evidence to support Darwin's theory of evolution by unearthing the ancestors of the modern horse. In 1877, Marsh delivered a very influential speech before the annual meeting of the American Association for the Advancement of Science, providing a demonstrative argument for evolution. For the first time, Marsh traced the evolution of vertebrates from fish all the way through humans. Sparing no detail, he listed a wealth of fossil examples of past life forms. The significance of this speech was immediately recognized by the scientific community, and it was printed in its entirety in several scientific journals.

Descent of Man, sexual selection, and botany

Further information: Orchids to Variation, Descent of Man to Emotions, and Insectivorous Plants to Worms

Despite repeated bouts of illness during the last twenty-two years of his life, Darwin's work continued. Having published On the Origin of Species as an abstract of his theory, he pressed on with experiments, research, and writing of his "big book". He covered human descent from earlier animals, including the evolution of society and of mental abilities, as well as explaining decorative beauty in wildlife and diversifying into innovative plant studies.

Enquiries about insect pollination led in 1861 to novel studies of wild orchids, showing adaptation of their flowers to attract specific moths to each species and ensure cross fertilisation. In 1862 Fertilisation of Orchids gave his first detailed demonstration of the power of natural selection to explain complex ecological relationships, making testable predictions. As his health declined, he lay on his sickbed in a room filled with inventive experiments to trace the movements of climbing plants. Admiring visitors included Ernst Haeckel, a zealous proponent of Darwinism incorporating Lamarckism and Goethe's idealism. Wallace remained supportive, though he increasingly turned to Spiritualism.

Darwin's book The Variation of Animals and Plants Under Domestication (1868) was the first part of his planned "big book", and included his unsuccessful hypothesis of pangenesis attempting to explain heredity. It sold briskly at first, despite its size, and was translated into many languages. He wrote most of a second part, on natural selection, but it remained unpublished in his lifetime.

Lyell had already popularised human prehistory, and Huxley had shown that anatomically humans are apes. With The Descent of Man, and Selection in Relation to Sex published in 1871, Darwin set out evidence from numerous sources that humans are animals, showing continuity of physical and mental attributes, and presented sexual selection to explain impractical animal features such as the peacock's plumage as well as human evolution of culture, differences between sexes, and physical and cultural racial classification, while emphasising that humans are all one species. According to an editorial in Nature journal: "Although Charles Darwin opposed slavery and proposed that humans have a common ancestor, he also advocated a hierarchy of races, with white people higher than others."

Letter of enquiry from Charles Darwin to the physiologist John Burdon-SandersonPunch's almanac for 1882, published shortly before Darwin's death, depicts him amidst evolution from chaos to Victorian gentleman with the title Man Is But A Worm

His research using images was expanded in his 1872 book The Expression of the Emotions in Man and Animals, one of the first books to feature printed photographs, which discussed the evolution of human psychology and its continuity with the behaviour of animals. Both books proved very popular, and Darwin was impressed by the general assent with which his views had been received, remarking that "everybody is talking about it without being shocked." His conclusion was "that man with all his noble qualities, with sympathy which feels for the most debased, with benevolence which extends not only to other men but to the humblest living creature, with his god-like intellect which has penetrated into the movements and constitution of the solar system – with all these exalted powers – Man still bears in his bodily frame the indelible stamp of his lowly origin."

His evolution-related experiments and investigations led to books on Insectivorous Plants, The Effects of Cross and Self Fertilisation in the Vegetable Kingdom, different forms of flowers on plants of the same species, and The Power of Movement in Plants. He continued to collect information and exchange views from scientific correspondents all over the world, including Mary Treat, whom he encouraged to persevere in her scientific work. He was the first person to recognise the significance of carnivory in plants. His botanical work was interpreted and popularised by various writers including Grant Allen and H. G. Wells, and helped transform plant science in the late 19th century and early 20th century.

Death and funeral

See also: Darwin from Insectivorous Plants to Worms § Death

In 1882, he was diagnosed with what was called "angina pectoris" which then meant coronary thrombosis and disease of the heart. At the time of his death, the physicians diagnosed "anginal attacks", and "heart-failure"; there has since been scholarly speculation about his life-long health issues.

He died at Down House on 19 April 1882. His last words were to his family, telling Emma, "I am not the least afraid of death—Remember what a good wife you have been to me—Tell all my children to remember how good they have been to me". While she rested, he repeatedly told Henrietta and Francis, "It's almost worthwhile to be sick to be nursed by you".

He had expected to be buried in St Mary's churchyard at Downe, but at the request of Darwin's colleagues, after public and parliamentary petitioning, William Spottiswoode (President of the Royal Society) arranged for Darwin to be honoured by burial in Westminster Abbey, close to John Herschel and Isaac Newton. The funeral, held on Wednesday 26 April, was attended by thousands of people, including family, friends, scientists, philosophers and dignitaries.

Children

Further information: Darwin–Wedgwood family § Charles Darwin

William Erasmus Darwin	27 December 1839 –	8 September 1914
Anne Elizabeth Darwin	2 March 1841 –	23 April 1851
Mary Eleanor Darwin	23 September 1842 –	16 October 1842
Henrietta Emma Darwin	25 September 1843 –	17 December 1927
George Howard Darwin	9 July 1845 –	7 December 1912
Elizabeth Darwin	8 July 1847 –	8 June 1926
Francis Darwin	16 August 1848 –	19 September 1925
Leonard Darwin	15 January 1850 –	26 March 1943
Horace Darwin	13 May 1851 –	29 September 1928
Charles Waring Darwin	6 December 1856 –	28 June 1858

The Darwins had ten children: two died in infancy, and Annie's death at the age of ten had a devastating effect on her parents. Charles was a devoted father and uncommonly attentive to his children. Whenever they fell ill, he feared that they might have inherited weaknesses from inbreeding due to the close family ties he shared with his wife and cousin, Emma Wedgwood. He examined inbreeding in his writings, contrasting it with the advantages of outcrossing in many species.

Charles Waring Darwin, born in December 1856, was the tenth and last of the children. Emma Darwin was aged 48 at the time of the birth, and the child was mentally subnormal and never learnt to walk or talk. He probably had Down syndrome, which had not then been medically described. The evidence is a photograph by William Erasmus Darwin of the infant and his mother, showing a characteristic head shape, and the family's observations of the child. Charles Waring died of scarlet fever on 28 June 1858, when Darwin wrote in his journal: "Poor dear Baby died."

Of his surviving children, George, Francis and Horace became Fellows of the Royal Society, distinguished as an astronomer, botanist and civil engineer, respectively. All three were knighted. Another son, Leonard, went on to be a soldier, politician, economist, eugenicist, and mentor of the statistician and evolutionary biologist Ronald Fisher.

Views and opinions

Religious views

Further information: Religious views of Charles Darwin

Darwin's family tradition was nonconformist Unitarianism, while his father and grandfather were freethinkers, and his baptism and boarding school were Church of England. When going to Cambridge to become an Anglican clergyman, he did not "in the least doubt the strict and literal truth of every word in the Bible". He learned John Herschel's science which, like William Paley's natural theology, sought explanations in laws of nature rather than miracles and saw adaptation of species as evidence of design. On board HMS Beagle, Darwin was quite orthodox and would quote the Bible as an authority on morality. He looked for "centres of creation" to explain distribution, and suggested that the very similar antlions found in Australia and England were evidence of a divine hand.

Upon his return, he expressed a critical view of the Bible's historical accuracy and questioned the basis for considering one religion more valid than another. In the next few years, while intensively speculating on geology and the transmutation of species, he gave much thought to religion and openly discussed this with his wife Emma, whose beliefs similarly came from intensive study and questioning.

The theodicy of Paley and Thomas Malthus vindicated evils such as starvation as a result of a benevolent creator's laws, which had an overall good effect. To Darwin, natural selection produced the good of adaptation but removed the need for design, and he could not see the work of an omnipotent deity in all the pain and suffering, such as the ichneumon wasp paralysing caterpillars as live food for its eggs. Though he thought of religion as a tribal survival strategy, Darwin was reluctant to give up the idea of God as an ultimate lawgiver. He was increasingly troubled by the problem of evil.

Darwin remained close friends with the vicar of Downe, John Brodie Innes, and continued to play a leading part in the parish work of the church, but from c. 1849 would go for a walk on Sundays while his family attended church. He considered it "absurd to doubt that a man might be an ardent theist and an evolutionist" and, though reticent about his religious views, in 1879 he wrote that "I have never been an atheist in the sense of denying the existence of a God. – I think that generally ... an agnostic would be the most correct description of my state of mind".

The "Lady Hope Story", published in 1915, claimed that Darwin had reverted to Christianity on his sickbed. The claims were repudiated by Darwin's children and have been dismissed as false by historians.

Human society

Darwin's views on social and political issues reflected his time and social position. He grew up in a family of Whig reformers who, like his uncle Josiah Wedgwood, supported electoral reform and the emancipation of slaves. Darwin was passionately opposed to slavery, while seeing no problem with the working conditions of English factory workers or servants.

Taking taxidermy lessons in 1826 from the freed slave John Edmonstone, whom Darwin long recalled as "a very pleasant and intelligent man", reinforced his belief that black people shared the same feelings, and could be as intelligent as people of other races. He took the same attitude to native people he met on the Beagle voyage. Though commonplace in Britain at the time, Silliman and Bachman noticed the contrast with slave-owning America. Around twenty years later, racism became a feature of British society, but Darwin remained strongly against slavery, against "ranking the so-called races of man as distinct species", and against ill-treatment of native people.

Darwin's interaction with Yaghans (Fuegians) such as Jemmy Button during the second voyage of HMS Beagle had a profound impact on his view of indigenous peoples. At his arrival in Tierra del Fuego he made a colourful description of "Fuegian savages". This view changed as he came to know Yaghan people more in detail. By studying the Yaghans, Darwin concluded that a number of basic emotions by different human groups were the same and that mental capabilities were roughly the same as for Europeans. While interested in Yaghan culture, Darwin failed to appreciate their deep ecological knowledge and elaborate cosmology until the 1850s when he inspected a dictionary of Yaghan detailing 32,000 words. He saw that European colonisation would often lead to the extinction of native civilisations, and "tr to integrate colonialism into an evolutionary history of civilization analogous to natural history".

Darwin's view of women was that men's eminence over them was the outcome of sexual selection, a view disputed by Antoinette Brown Blackwell in her 1875 book The Sexes Throughout Nature.

Darwin was intrigued by his half-cousin Francis Galton's argument, introduced in 1865, that statistical analysis of heredity showed that moral and mental human traits could be inherited, and principles of animal breeding could apply to humans. In The Descent of Man, Darwin noted that aiding the weak to survive and have families could lose the benefits of natural selection, but cautioned that withholding such aid would endanger the instinct of sympathy, "the noblest part of our nature", and factors such as education could be more important. When Galton suggested that publishing research could encourage intermarriage within a "caste" of "those who are naturally gifted", Darwin foresaw practical difficulties and thought it "the sole feasible, yet I fear utopian, plan of procedure in improving the human race", preferring to simply publicise the importance of inheritance and leave decisions to individuals. Francis Galton named this field of study "eugenics" in 1883, after Darwin's death, and his theories were cited to promote eugenic policies.

Evolutionary social movements

Further information: Eugenics, Social effects of evolutionary theory, and Degeneration theory

Darwin's fame and popularity led to his name being associated with ideas and movements that, at times, had only an indirect relation to his writings, and sometimes went directly against his express comments.

Thomas Malthus had argued that population growth beyond resources was ordained by God to get humans to work productively and show restraint in getting families; this was used in the 1830s to justify workhouses and laissez-faire economics. Evolution was by then seen as having social implications, and Herbert Spencer's 1851 book Social Statics based ideas of human freedom and individual liberties on his Lamarckian evolutionary theory.

Soon after the Origin was published in 1859, critics derided his description of a struggle for existence as a Malthusian justification for the English industrial capitalism of the time. The term Darwinism was used for the evolutionary ideas of others, including Spencer's "survival of the fittest" as free-market progress, and Ernst Haeckel's polygenistic ideas of human development. Writers used natural selection to argue for various, often contradictory, ideologies such as laissez-faire dog-eat-dog capitalism, colonialism and imperialism. However, Darwin's holistic view of nature included "dependence of one being on another"; thus pacifists, socialists, liberal social reformers and anarchists such as Peter Kropotkin stressed the value of cooperation over struggle within a species. Darwin himself insisted that social policy should not simply be guided by concepts of struggle and selection in nature.

After the 1880s, a eugenics movement developed on ideas of biological inheritance, and for scientific justification of their ideas appealed to some concepts of Darwinism. In Britain, most shared Darwin's cautious views on voluntary improvement and sought to encourage those with good traits in "positive eugenics". During the "Eclipse of Darwinism", a scientific foundation for eugenics was provided by Mendelian genetics. Negative eugenics to remove the "feebleminded" were popular in America, Canada and Australia, and eugenics in the United States introduced compulsory sterilisation laws, followed by several other countries. Subsequently, Nazi eugenics brought the field into disrepute.

The term "Social Darwinism" was used infrequently from around the 1890s, but became popular as a derogatory term in the 1940s when used by Richard Hofstadter to attack the laissez-faire conservatism of those like William Graham Sumner who opposed reform and socialism. Since then, it has been used as a term of abuse by those opposed to what they think are the moral consequences of evolution.

Works

Further information: Charles Darwin bibliography

Darwin was a prolific writer. Even without the publication of his works on evolution, he would have had a considerable reputation as the author of The Voyage of the Beagle, as a geologist who had published extensively on South America and had solved the puzzle of the formation of coral atolls, and as a biologist who had published the definitive work on barnacles. While On the Origin of Species dominates perceptions of his work, The Descent of Man and The Expression of the Emotions in Man and Animals had considerable impact, and his books on plants including The Power of Movement in Plants were innovative studies of great importance, as was his final work on The Formation of Vegetable Mould through the Action of Worms.

Legacy and commemoration

Further information: List of things named after Charles Darwin, List of taxa described by Charles Darwin, and Commemoration of Charles Darwin

As Alfred Russel Wallace put it, Darwin had "wrought a greater revolution in human thought within a quarter of a century than any man of our time – or perhaps any time", having "given us a new conception of the world of life, and a theory which is itself a powerful instrument of research; has shown us how to combine into one consistent whole the facts accumulated by all the separate classes of workers, and has thereby revolutionised the whole study of nature". The paleoanthropologist Trenton Holliday states that "Darwin is rightly considered to be the preeminent evolutionary scientist of all time".

By around 1880, most scientists were convinced of evolution as descent with modification, though few agreed with Darwin that natural selection "has been the main but not the exclusive means of modification". During "the eclipse of Darwinism" scientists explored alternative mechanisms. Then Ronald Fisher incorporated Mendelian genetics in The Genetical Theory of Natural Selection, leading to population genetics and the modern evolutionary synthesis, which continues to develop. Scientific discoveries have confirmed and validated Darwin's key insights.

Geographical features given his name include Darwin Sound and Mount Darwin, both named while he was on the Beagle voyage, and Darwin Harbour, named by his former shipmates on its next voyage, which eventually became the location of Darwin, the capital city of Australia's Northern Territory. Darwin's name was given, formally or informally, to numerous plants and animals, including many he had collected on the voyage. The Linnean Society of London began awards of the Darwin–Wallace Medal in 1908, to mark fifty years from the joint reading on 1 July 1858 of papers by Darwin and Wallace publishing their theory. Further awards were made in 1958 and 2008; since 2010, the awards have been annual. Darwin College, a postgraduate college at Cambridge University founded in 1964, is named after the Darwin family. From 2000 to 2017, UK £10 banknotes issued by the Bank of England featured Darwin's portrait printed on the reverse, along with a hummingbird and HMS Beagle.

See also

• 1991 Darwin
• Creation (biographical drama film)
• Creation–evolution controversy
• European and American voyages of scientific exploration
• History of biology
• History of evolutionary thought
• List of coupled cousins
• List of multiple discoveries
• Multiple discovery
• Portraits of Charles Darwin
• Tinamou egg
• Universal Darwinism

Notes

I. Robert FitzRoy was to become known after the voyage for biblical literalism, but at this time he had considerable interest in Lyell's ideas, and they met before the voyage when Lyell asked for observations to be made in South America. FitzRoy's diary during the ascent of the River Santa Cruz in Patagonia recorded his opinion that the plains were raised beaches, but on return, newly married to a very religious lady, he recanted these ideas.(Browne 1995, pp. 186, 414)

II. In the section "Morphology" of Chapter XIII of On the Origin of Species, Darwin commented on homologous bone patterns between humans and other mammals, writing: "What can be more curious than that the hand of a man, formed for grasping, that of a mole for digging, the leg of the horse, the paddle of the porpoise, and the wing of the bat, should all be constructed on the same pattern, and should include the same bones, in the same relative positions?" and in the concluding chapter: "The framework of bones being the same in the hand of a man, wing of a bat, fin of the porpoise, and leg of the horse … at once explain themselves on the theory of descent with slow and slight successive modifications."

III. In On the Origin of Species Darwin mentioned human origins in his concluding remark that "In the distant future I see open fields for far more important researches. Psychology will be based on a new foundation, that of the necessary acquirement of each mental power and capacity by gradation. Light will be thrown on the origin of man and his history."

In "Chapter VI: Difficulties on Theory" he referred to sexual selection: "I might have adduced for this same purpose the differences between the races of man, which are so strongly marked; I may add that some little light can apparently be thrown on the origin of these differences, chiefly through sexual selection of a particular kind, but without here entering on copious details my reasoning would appear frivolous."

In The Descent of Man of 1871, Darwin discussed the first passage: "During many years I collected notes on the origin or descent of man, without any intention of publishing on the subject, but rather with the determination not to publish, as I thought that I should thus only add to the prejudices against my views. It seemed to me sufficient to indicate, in the first edition of my 'Origin of Species,' that by this work 'light would be thrown on the origin of man and his history;' and this implies that man must be included with other organic beings in any general conclusion respecting his manner of appearance on this earth." In a preface to the 1874 second edition, he added a reference to the second point: "it has been said by several critics, that when I found that many details of structure in man could not be explained through natural selection, I invented sexual selection; I gave, however, a tolerably clear sketch of this principle in the first edition of the 'Origin of Species,' and I there stated that it was applicable to man."

IV. See, for example, WILLA volume 4, Charlotte Perkins Gilman and the Feminization of Education by Deborah M. De Simone: "Gilman shared many basic educational ideas with the generation of thinkers who matured during the period of "intellectual chaos" caused by Darwin's Origin of the Species. Marked by the belief that individuals can direct human and social evolution, many progressives came to view education as the panacea for advancing social progress and for solving such problems as urbanisation, poverty, or immigration."

V. See, for example, the song "A lady fair of lineage high" from Gilbert and Sullivan's Princess Ida, which describes the descent of man (but not woman!) from apes.

VI. Darwin's belief that black people had the same essential humanity as Europeans, and had many mental similarities, was reinforced by the lessons he had from John Edmonstone in 1826. Early in the Beagle voyage, Darwin nearly lost his position on the ship when he criticised FitzRoy's defence and praise of slavery. (Darwin 1958, p. 74) He wrote home about "how steadily the general feeling, as shown at elections, has been rising against Slavery. What a proud thing for England if she is the first European nation which utterly abolishes it! I was told before leaving England that after living in slave countries all my opinions would be altered; the only alteration I am aware of is forming a much higher estimate of the negro character." (Darwin 1887, p. 246) Regarding Fuegians, he "could not have believed how wide was the difference between savage and civilized man: it is greater than between a wild and domesticated animal, inasmuch as in man there is a greater power of improvement", but he knew and liked civilised Fuegians like Jemmy Button: "It seems yet wonderful to me, when I think over all his many good qualities, that he should have been of the same race, and doubtless partaken of the same character, with the miserable, degraded savages whom we first met here." (Darwin 1845, pp. 205, 207–208)

In the Descent of Man, he mentioned the similarity of Fuegians' and Edmonstone's minds to Europeans' when arguing against "ranking the so-called races of man as distinct species".

He rejected the ill-treatment of native people, and for example wrote of massacres of Patagonian men, women, and children, "Every one here is fully convinced that this is the most just war, because it is against barbarians. Who would believe in this age that such atrocities could be committed in a Christian civilized country?"(Darwin 1845, p. 102)

VII. Geneticists studied human heredity as Mendelian inheritance, while eugenics movements sought to manage society, with a focus on social class in the United Kingdom, and on disability and ethnicity in the United States, leading to geneticists seeing this movement as impractical pseudoscience. A shift from voluntary arrangements to "negative" eugenics included compulsory sterilisation laws in the United States, copied by Nazi Germany as the basis for Nazi eugenics based on virulent racism and "racial hygiene".
(Thurtle, Phillip (17 December 1996). "the creation of genetic identity". SEHR. Vol. 5, no. Supplement: Cultural and Technological Incubations of Fascism. Retrieved 11 November 2008. Edwards, A. W. F. (1 April 2000). "The Genetical Theory of Natural Selection". Genetics. Vol. 154, no. April 2000. pp. 1419–1426. PMC 1461012. PMID 10747041. Retrieved 11 November 2008.Wilkins, John. "Evolving Thoughts: Darwin and the Holocaust 3: eugenics". Archived from the original on 5 December 2008. Retrieved 11 November 2008.)

VIII. David Quammen writes of his "theory that turned to these arcane botanical studies – producing more than one book that was solidly empirical, discreetly evolutionary, yet a 'horrid bore' – at least partly so that the clamorous controversialists, fighting about apes and angels and souls, would leave him... alone". David Quammen, "The Brilliant Plodder" (review of Ken Thompson, Darwin's Most Wonderful Plants: A Tour of His Botanical Legacy, University of Chicago Press, 255 pp.; Elizabeth Hennessy, On the Backs of Tortoises: Darwin, the Galápagos, and the Fate of an Evolutionary Eden, Yale University Press, 310 pp.; Bill Jenkins, Evolution Before Darwin: Theories of the Transmutation of Species in Edinburgh, 1804–1834, Edinburgh University Press, 222 pp.), The New York Review of Books, vol. LXVII, no. 7 (23 April 2020), pp. 22–24. Quammen, quoted from p. 24 of his review.

Citations

1. Freeman 2007, p. 76.
2. "Charles Darwin's personal finances revealed in new find". The Telegraph. 22 March 2009. Archived from the original on 19 October 2017.
3. ^ "Search Results: Record – Darwin; Charles Robert". The Royal Society Collections Catalogues. 20 June 2015. Archived from the original on 2 December 2021. Retrieved 2 December 2021.
4. ^ Freeman 2007, p. 106.
5. "Darwin" Archived 18 July 2014 at the Wayback Machine entry in Collins English Dictionary.
6. Desmond, Moore & Browne 2004.
7. Coyne, Jerry A. (2009). Why Evolution is True. Viking. pp. 8–11. ISBN 978-0-670-02053-9.
8. Larson 2004, pp. 79–111.
9. "Special feature: Darwin 200". New Scientist. Archived from the original on 11 February 2011. Retrieved 2 April 2011.
10. ^ "Westminster Abbey » Charles Darwin". Westminster Abbey. 2 January 2016. Archived from the original on 4 March 2016. Retrieved 2 January 2016.
    Leff 2000, Darwin's Burial
11. ^ Leff 2000, About Charles Darwin.
12. Desmond & Moore 1991, pp. 210, 284–285.
13. Desmond & Moore 1991, pp. 263–274.
14. van Wyhe 2007, pp. 184, 187
15. Beddall, B. G. (1968). "Wallace, Darwin, and the Theory of Natural Selection". Journal of the History of Biology. 1 (2): 261–323. doi:10.1007/BF00351923. ISSN 0022-5010. S2CID 81107747.
16. ^ van Wyhe 2008.
17. Coyne, Jerry A. (2009). Why Evolution is True. Oxford: Oxford University Press. p. 17. ISBN 978-0-19-923084-6. In The Origin, Darwin provided an alternative hypothesis for the development, diversification, and design of life. Much of that book presents evidence that not only supports evolution but at the same time refutes creationism. In Darwin's day, the evidence for his theories was compelling but not completely decisive.
18. Glass, Bentley (1959). Forerunners of Darwin. Baltimore, MD: Johns Hopkins University Press. p. iv. ISBN 978-0-8018-0222-5. Darwin's solution is a magnificent synthesis of evidence ... a synthesis ... compelling in honesty and comprehensiveness.
19. ^ Bowler 2003, pp. 178–179, 338, 347.
20. Desmond, Adrian J. (13 September 2002). "Charles Darwin". Encyclopædia Britannica. Archived from the original on 6 February 2018. Retrieved 11 February 2018.
21. John H. Wahlert (11 June 2001). "The Mount House, Shrewsbury, England (Charles Darwin)". Darwin and Darwinism. Baruch College. Archived from the original on 6 December 2008. Retrieved 26 November 2008.
22. Smith, Homer W. (1952). Man and His Gods. New York: Grosset & Dunlap. pp. 339–340.
23. ^ Desmond & Moore 1991, pp. 12–15;
    Darwin 1958, pp. 21–25.
24. Darwin 1958, pp. 46–48.
25. ^ Darwin 1958, p. 51;
    Desmond & Moore 2009, pp. 18–26.
26. Desmond & Moore 1991, pp. 31–34.
27. Browne 1995, pp. 72–88.
28. Desmond & Moore 1991, pp. 42–43.
29. Browne 1995, pp. 47–48, 89–91;
    Desmond & Moore 2009, pp. 47–48.
30. Darwin 1887, p. 48.
31. "Darwin statue unveiled at college". BBC News. 12 February 2009. Archived from the original on 16 February 2009. Retrieved 22 April 2022.
32. ^ Smith, Homer W. (1952). Man and His Gods. New York: Grosset & Dunlap. pp. 357–358.
33. ^ Darwin 1887, pp. 50–51.
34. van Wyhe, John (ed.). "Darwin's insects in Stephens' Illustrations of British entomology (1829–32)". Darwin Online. Archived from the original on 1 September 2019. Retrieved 3 July 2020.
35. ^ Desmond & Moore 1991, pp. 73–79, 763;
    Darwin 1958, pp. 57–67.
36. Browne 1995, p. 97.
37. ^ von Sydow 2005, pp. 5–7.
38. Daum, Andreas W. (2024). Alexander von Humboldt: A Concise Biography. Trans. Robert Savage. Princeton, N.J.: Princeton University Press. pp. 137–138. ISBN 978-0-691-24736-6.
39. ^ Darwin 1958, pp. 67–68.
40. Browne 1995, pp. 128–129, 133–141.
41. Peter Lucas (1 January 2010). "The recovery of time past: Darwin at Barmouth on the eve of the Beagle". Darwin Online. Archived from the original on 11 April 2010. Retrieved 5 December 2021.
42. "Letter no. 105, Henslow, J. S. to Darwin, C. R., 24 Aug 1831". Darwin Correspondence Project. Archived from the original on 29 December 2021. Retrieved 29 December 2021.
43. Desmond & Moore 1991, pp. 94–97
44. Browne 1995, pp. 204–210
45. ^ Keynes 2000, pp. ix–xi
46. van Wyhe 2008b, pp. 18–21
47. Gordon Chancellor; Randal Keynes (October 2006). "Darwin's field notes on the Galapagos: 'A little world within itself'". Darwin Online. Archived from the original on 1 September 2009. Retrieved 16 September 2009.
48. Keynes 2001, pp. 21–22
49. Browne 1995, pp. 183–190
50. Keynes 2001, pp. 41–42
51. Darwin 1958, pp. 73–74
52. Browne 1995, pp. 223–225
    Darwin 1835, p. 7
    "Letter no. 213, Henslow, J. S. to Darwin, C. R., 31 August 1833". Darwin Correspondence Project. Archived from the original on 29 December 2021. Retrieved 29 December 2021.
53. ^ Keynes 2001, pp. 106–109
54. Jaksic, Fabian M. (2022). "Historical account and current ecological knowledge of the southernmost lizard in the world, Liolaemus magellanicus (Squamata: Liolaemidae)". Revista Chilena de Historia Natural. 95 (7). Bibcode:2022RvCHN..95....7J. doi:10.1186/s40693-022-00112-y. S2CID 252717680.
55. Desmond & Moore 1991, pp. 189–192, 198
56. Eldredge 2006
57. Desmond & Moore 1991, pp. 131, 159
    Herbert 1991, pp. 174–179
58. "Darwin Online: 'Hurrah Chiloe': an introduction to the Port Desire Notebook". Archived from the original on 4 December 2008. Retrieved 24 October 2008.
59. Darwin 1845, pp. 205–208
60. Browne 1995, pp. 243–244, 248–250, 382–383
61. Keynes 2001, pp. 226–227
62. Desmond & Moore 1991, pp. 160–168, 182
    "Letter no. 275 – Charles Darwin to Susan Elizabeth Darwin – 23 April 1835". Darwin Correspondence Project. Archived from the original on 6 December 2021. Retrieved 6 December 2021.
63. ^ Darwin 1958, pp. 98–99
64. ^ Keynes 2001, pp. 356–357
65. Sulloway 1982, p. 19
66. ^ "Darwin Online: Coccatoos & Crows: An introduction to the Sydney Notebook". Archived from the original on 14 January 2009. Retrieved 2 January 2009.
67. Keynes 2001, pp. 398–399.
68. "Letter no. 301, Charles Darwin to Caroline Darwin, 29 April 1836, Port Lewis, Mauritius". Darwin Correspondence Project. Archived from the original on 15 February 2022. Retrieved 12 February 2022.
69. Browne 1995, p. 336
70. ^ Darwin 1839, p. viii
71. ^ van Wyhe 2007, p. 197
72. Keynes 2000, pp. xix–xx
    Eldredge 2006
73. Darwin 1859, p. 1
74. Darwin 1835, p. 1.
75. "Letter no. 291, Caroline Darwin to Charles Darwin, 29 December [1835], [Shrewsbury]". Darwin Correspondence Project. Archived from the original on 11 February 2022. Retrieved 19 January 2022.
76. "Letter no. 302, Charles Darwin to Catherine Darwin, 3 June 1836, Cape of Good Hope". Darwin Correspondence Project. Archived from the original on 27 January 2022. Retrieved 19 January 2022.
77. "Letter no. 288, Susan Darwin to Charles Darwin, 22 November 1835, Shrewsbury". Darwin Correspondence Project. Archived from the original on 13 February 2022. Retrieved 19 January 2022.
78. Darwin 1958, pp. 81–82.
79. Desmond & Moore 1991, pp. 195–198
80. Owen 1840, pp. 16, 73, 106
    Eldredge 2006
81. Desmond & Moore 1991, pp. 201–205
    Browne 1995, pp. 349–350
82. Browne 1995, pp. 345–347.
83. Keynes 2001, pp. xviii–xix.
84. Desmond & Moore 1991, pp. 207–210
    Sulloway 1982, pp. 20–23
85. "Darwin Correspondence Project – Letter 346 – Darwin, C. R. to Darwin, C. S., 27 Feb 1837". Archived from the original on 29 June 2009. Retrieved 19 December 2008. proposes a move on Friday 3 March 1837,
    Darwin's Journal (Darwin 2006, pp. 12 verso) backdated from August 1838 gives a date of 6 March 1837
86. Desmond & Moore 1991, pp. 201, 212–221
87. Sulloway 1982, pp. 9, 20–23
88. Browne 1995, p. 360
    "Darwin, C. R. (Read 14 March 1837) Notes on Rhea americana and Rhea darwinii, Proceedings of the Zoological Society of London". Archived from the original on 10 February 2009. Retrieved 17 December 2008.
89. Herbert 1980, pp. 7–10
    van Wyhe 2008b, p. 44
    Darwin 1837, pp. 1–13, 26, 36, 74
    Desmond & Moore 1991, pp. 229–232
90. "£1,000 in 1832 → 2021 | UK Inflation Calculator". www.in2013dollars.com. Archived from the original on 15 August 2021. Retrieved 8 August 2021.
91. Browne 1995, pp. 367–369
92. ^ Keynes 2001, p. xix
93. Desmond & Moore 1991, pp. 233–234
    "Darwin Correspondence Project – Letter 404 – Buckland, William to Geological Society of London, 9 Mar 1838". Archived from the original on 29 June 2009. Retrieved 23 December 2008.
94. Desmond & Moore 1991, pp. 233–236.
95. Desmond & Moore 1991, pp. 241–244, 426
96. Browne 1995, p. xii
97. Desmond & Moore 1991, pp. 241–244
98. Desmond & Moore 1991, pp. 252, 476, 531
    Darwin 1958, p. 115
99. Desmond & Moore 1991, p. 254
    Browne 1995, pp. 377–378
    Darwin 1958, p. 84
100. Darwin 1958, pp. 232–233
101. Desmond & Moore 1991, pp. 256–259
102. "Darwin transmutation notebook D pp. 134e–135e". Archived from the original on 18 July 2012. Retrieved 4 June 2012.
103. Desmond & Moore 1991, pp. 264–265
     Browne 1995, pp. 385–388
     Darwin 1842, p. 7
104. ^ Darwin 1958, p. 120
105. "Darwin transmutation notebook E p. 75". Archived from the original on 28 June 2009. Retrieved 17 March 2009.
106. "Darwin transmutation notebook E p. 71". Archived from the original on 28 June 2009. Retrieved 17 March 2009.
107. ^ "Darwin Correspondence Project – Belief: historical essay". Archived from the original on 25 February 2009. Retrieved 25 November 2008.
108. Desmond & Moore 1991, pp. 272–279
109. Desmond & Moore 1991, p. 279
110. "Darwin Correspondence Project – Letter 419 – Darwin, C. R. to Fox, W. D., (15 June 1838)". Darwin Correspondence Project. Archived from the original on 4 September 2007. Retrieved 8 February 2008.
111. van Wyhe 2007, pp. 186–192
112. Buchanan, Roderick D. (May 2017). "Darwin's "Mr. Arthrobalanus": Sexual Differentiation, Evolutionary Destiny and the Expert Eye of the Beholder". Journal of the History of Biology. 50 (2): 315–355. doi:10.1007/s10739-016-9444-9. ISSN 1573-0387. PMID 27098777. Retrieved 3 September 2024.
113. Desmond & Moore 1991, pp. 284–285, 292
114. Desmond & Moore 1991, pp. 292–293
     Darwin 1842, pp. xvi–xvii
115. Darwin 1958, p. 114
116. van Wyhe 2007, pp. 183–184
117. "Darwin Correspondence Project – Letter 729 – Darwin, C. R. to Hooker, J. D., (11 January 1844)". Archived from the original on 7 March 2008. Retrieved 8 February 2008.
118. "Darwin Correspondence Project – Letter 734 – Hooker, J. D. to Darwin, C. R., 29 January 1844". Archived from the original on 26 February 2009. Retrieved 8 February 2008.
119. "Charles Darwin: a life in pictures, The Sand Walk near Down House, Darwin's thinking path". Darwin Online. Archived from the original on 1 October 2022. Retrieved 1 October 2022.
120. van Wyhe 2007, p. 188
121. Browne 1995, pp. 461–465
122. "Darwin Correspondence Project – Letter 814 – Darwin, C. R. to Hooker, J. D., (7 Jan 1845)". Archived from the original on 5 December 2008. Retrieved 24 November 2008.
123. van Wyhe 2007, pp. 190–191
124. Desmond & Moore 1991, pp. 320–323, 339–348
125. "Darwin Correspondence Project – Letter 1236 – Darwin, C. R. to Hooker, J. D., 28 Mar 1849". Archived from the original on 7 December 2008. Retrieved 24 November 2008.
126. Browne 1995, pp. 498–501
127. Darwin 1958, pp. 117–118
128. Desmond & Moore 1991, pp. 383–387
129. Bromham, Lindell (1 October 2020). "Comparability in evolutionary biology: The case of Darwin's barnacles". Linguistic Typology. 24 (3): 427–463. doi:10.1515/lingty-2020-2056. hdl:1885/274303. ISSN 1613-415X. S2CID 222319487.
130. van Wyhe 2007.
131. Freeman 2007, pp. 107, 109
132. Desmond & Moore 1991, pp. 419–420
133. Darwin Online: Photograph of Charles Darwin by Maull and Polyblank for the Literary and Scientific Portrait Club (1855) Archived 7 January 2012 at the Wayback Machine, John van Wyhe, December 2006
134. ^ Desmond & Moore 1991, pp. 412–441, 457–458, 462–463
     Desmond & Moore 2009, pp. 283–284, 290–292, 295
135. Ball, P. (2011). Shipping timetables debunk Darwin plagiarism accusations: Evidence challenges claims that Charles Darwin stole ideas from Alfred Russel Wallace. Nature. online Archived 22 February 2012 at the Wayback Machine
136. van Wyhe, John; Rookmaaker, Kees (2012). "A new theory to explain the receipt of Wallace's Ternate Essay by Darwin in 1858". Biological Journal of the Linnean Society. 105: 249–252. doi:10.1111/j.1095-8312.2011.01808.x.
137. Desmond & Moore 1991, pp. 466–470
138. Browne 2002, pp. 40–42, 48–49
139. Darwin 1958, p. 122
140. Desmond & Moore 1991, pp. 374–474
141. Desmond & Moore 1991, p. 477
142. Darwin 1859, p. 459
143. van Wyhe 2008.
144. ^ Darwin 1859, p. 199
     Darwin & Costa 2009, p. 199
     Desmond & Moore 2009, p. 310
145. ^ Darwin 1859, p. 488
     Darwin & Costa 2009, pp. 199, 488
     van Wyhe 2008
146. Darwin 1859, p. 5
147. Darwin 1859, p. 492
148. Browne 2002, p. 59, Freeman 1977, pp. 79–80
149. ^ Browne 2002, pp. 373–379
150. van Wyhe 2008b, p. 48
151. Browne 2002, pp. 103–104, 379
152. Radick 2013, pp. 174–175
     Huxley & Kettlewell 1965, p. 88
153. Browne 2002, p. 87
     Leifchild 1859
154. Desmond & Moore 1991, pp. 477–491
155. Browne 2002, pp. 110–112
156. Bowler 2003, pp. 158, 186
157. ^ "Darwin and design: historical essay". Darwin Correspondence Project. 2007. Archived from the original on 15 June 2009. Retrieved 17 September 2008.
158. Desmond & Moore 1991, pp. 487–488, 500
159. ^ Miles 2001
160. Bowler 2003, p. 185
161. Browne 2002, pp. 156–159
162. "Science ahead of its time: Secret of 157-year old Darwin manuscript". National University of Singapore News. 24 November 2022. Archived from the original on 25 November 2022. Retrieved 25 November 2022.
163. Peretó, Juli; Bada, Jeffrey L.; Lazcano, Antonio (1 October 2009). "Charles Darwin and the Origin of Life". Origins of Life and Evolution of Biospheres. 39 (5): 395–406. doi:10.1007/s11084-009-9172-7. ISSN 1573-0875. PMC 2745620. PMID 19633921. Archived from the original on 18 December 2023. Retrieved 8 December 2023.
164. ^ Browne 2002, pp. 217–226
165. "Darwin Correspondence Project – Letter 4652 – Falconer, Hugh to Darwin, C. R., 3 Nov (1864)". Archived from the original on 5 December 2008. Retrieved 1 December 2008.
166. "Darwin Correspondence Project – Letter 4807 – Hooker, J. D. to Darwin, C. R., (7–8 Apr 1865)". Archived from the original on 5 December 2008. Retrieved 1 December 2008.
167. Bowler 2003, p. 196
168. Desmond & Moore 1991, pp. 507–508
     Browne 2002, pp. 128–129, 138
169. Plate, Robert. The Dinosaur Hunters: Othniel C. Marsh and Edward D. Cope, pp. 69, 203-5, David McKay Company, Inc., New York, 1964.
170. McCarren, Mark J. The Scientific Contributions of Othniel Charles Marsh, pp. 37-9, Peabody Museum of Natural History, Yale University, New Haven, Connecticut, 1993. ISBN 0-912532-32-7
171. Plate, Robert. The Dinosaur Hunters: Othniel C. Marsh and Edward D. Cope, pp. 188-9, David McKay Company, Inc., New York, 1964.
172. Desmond, Moore & Browne 2007, pp. 73–75.
173. Desmond, Moore & Browne 2007, pp. 78–83, 86–90.
174. van Wyhe 2008b, pp. 50–55
175. "The correspondence of Charles Darwin, volume 14: 1866". Archived from the original on 5 June 2010. Retrieved 6 March 2009. Cambridge University Press. Retrieved 25 June 2012
176. Smith 1999.
177. Freeman 1977, p. 122
178. Darwin 1871, pp. 385–405
     Browne 2002, pp. 339–343
179. Nobles, Melissa; Womack, Chad; Wonkam, Ambroise; Wathuti, Elizabeth (8 June 2022). "Science must overcome its racist legacy: Nature's guest editors speak". Nature. 606 (7913): 225–227. Bibcode:2022Natur.606..225N. doi:10.1038/d41586-022-01527-z. PMID 35676434. S2CID 249520597. "In The Descent of Man, Darwin describes what he calls the gradations between the highest men of the highest races and the lowest savages*. He uses the word 'savages' to describe Black and Indigenous people." (*see Darwin, C. R. The Descent of Man, and Selection in Relation to Sex (John Murray, 1871))
180. Browne 2002, pp. 359–369
     Darwin 1887, p. 133
181. Darwin 1871, p. 405
182. Darwin's Women Archived 12 February 2020 at the Wayback Machine at Cambridge University
183. Hedrich, Rainer; Fukushima, Kenji (17 June 2021). "On the Origin of Carnivory: Molecular Physiology and Evolution of Plants on an Animal Diet". Annual Review of Plant Biology. 72 (1): 133–153. doi:10.1146/annurev-arplant-080620-010429. ISSN 1543-5008. PMID 33434053. S2CID 231595236.
184. Pain, Stephanie (2 March 2022). "How plants turned predator". Knowable Magazine. doi:10.1146/knowable-030122-1. Archived from the original on 8 March 2022. Retrieved 11 March 2022.
185. Endersby, Jim (June 2016). "Deceived by orchids: sex, science, fiction and Darwin". The British Journal for the History of Science. 49 (2): 205–229. doi:10.1017/S0007087416000352. ISSN 0007-0874. PMID 27278105. S2CID 23027055. Retrieved 17 March 2022.
186. Colp, Ralph (2008). "The Final Illnes [sic]". Darwin's Illness. pp. 116–120. doi:10.5744/florida/9780813032313.003.0014. ISBN 978-0-8130-3231-3.
187. Clayton, Julie (24 June 2010). "Chagas disease 101". Nature. 465 (n7301_supp): S4 – S5. Bibcode:2010Natur.465S...3C. doi:10.1038/nature09220. PMID 20571553. S2CID 205221512.
188. Darwin, Emma (1882). "[Reminiscences of Charles Darwin's last years.] CUL-DAR210.9". Archived from the original on 28 June 2009. Retrieved 8 January 2009.
189. Desmond & Moore 1991, pp. 664–677
190. Desmond & Moore 1991, p. 447.
191. David P. Steensma (15 March 2005). "Down syndrome in Down House: trisomy 21, GATA1 mutations, and Charles Darwin". Blood 105 (6) 2614–2616.
192. Freeman, R. B. (1984), Darwin Pedigrees, London, p. 43.
193. Darwin, C. R. Journal (1809–1881), p. 37.
194. "List of Fellows of the Royal Society, 1660–2006, A–J". Archived from the original (PDF) on 3 February 2017. Retrieved 16 September 2009.
195. O'Connor, John J.; Robertson, Edmund F., "Charles Darwin", MacTutor History of Mathematics Archive, University of St Andrews
196. Berra, Tim M. Darwin and His Children: His Other Legacy, (Oxford: 2013, Oxford UP), 101, 129, 168. George became a knight commander of the Order of the Bath in 1905. Francis was knighted in 1912. Horace became a knight commander of the KBE in 1918.
197. Edwards, A. W. F. 2004. Darwin, Leonard (1850–1943). In: Oxford Dictionary of National Biography, Oxford University Press.
198. ^ Darwin 1958, pp. 85–96
199. ^ van Wyhe 2008b, p. 41
200. von Sydow 2005, pp. 8–14
201. von Sydow 2005, pp. 4–5, 12–14
202. Moore 2006
203. "Darwin Correspondence Project – Darwin and the church: historical essay". 5 June 2015. Archived from the original on 28 November 2016. Retrieved 26 November 2016.
204. ^ Letter 12041 Archived 7 November 2009 at the Wayback Machine – Darwin, C. R. to Fordyce, John, 7 May 1879
205. Darwin's Complex loss of Faith Archived 11 February 2017 at the Wayback Machine The Guardian 17 September 2009
206. Moore 2005
     Yates 2003
207. Browne 1995, pp. 196–197.
208. Browne 1995, pp. 66, 198, 240.
209. Silliman, B. (1810). A Journal of Travels in England, Holland and Scotland: And of Two Passages Over the Atlantic, in the Years 1805 and 1806 ... D. & G. Bruce. pp. 216–217. Archived from the original on 18 December 2023. Retrieved 29 August 2022. As there are no slaves in England, perhaps the English have not learned to regard negroes as a degraded class of men, as we do in the United States
     Bachman, J. (1850). The Doctrine of the Unity of the Human Race Examined on the Principles of Science. American culture series. C. Canning. p. 105. ISBN 978-0-608-43507-7. Archived from the original on 18 September 2023. Retrieved 29 August 2022.
210. Wilkins 2008, pp. 408–413
211. ^ Rozzi, Ricardo (2018). "Transformaciones del pensamiento de Darwin en cabo de hornos: Un legado para la ciencia y la etica ambiental" [Transformations of Darwin's thought in cape horn: A legacy for science and environmental ethics]. Magallania (in Spanish). 46 (1): 267–277. doi:10.4067/S0718-22442018000100267.
212. ^ Barta, Tony (2 June 2005). "Mr Darwin's shooters: on natural selection and the naturalizing of genocide". Patterns of Prejudice. 39 (2): 116–137. doi:10.1080/00313220500106170. S2CID 159807728.
213. Vandermassen, Griet (2004). "Sexual Selection: A Tale of Male Bias and Feminist Denial". European Journal of Women's Studies. 11 (9): 11–13. CiteSeerX 10.1.1.550.3672. doi:10.1177/1350506804039812. S2CID 145221350.
214. Desmond & Moore 1991, pp. 556–557, 572, 598
     Darwin 1871, pp. 167–173, 402–403
     "Correspondence between Francis Galton and Charles Darwin". Archived from the original on 2 January 2009. Retrieved 8 November 2008.
215. ^ Wilkins 1997
     Moore 2006
216. Sweet 2004
217. Paul 2003, pp. 223–225
218. Bannister 1989
219. Paul 2003
     Kotzin 2004
220. Balfour 1882
     van Wyhe 2008
     Anonymous 1882
221. Brummitt, R. K.; C. E. Powell (1992). Authors of Plant Names. Royal Botanic Gardens, Kew. ISBN 978-1-84246-085-6.
222. ^ Van Wyhe, J. (2021). Charles Darwin: The Man, His Great Voyage, and His Theory of Evolution. Pioneers of Science. Rosen Publishing Group, Incorporated. pp. 154–155. ISBN 978-1-4994-7110-6. Archived from the original on 18 December 2023. Retrieved 23 May 2022.
223. Holliday 2021, p. 5.
224. van Wyhe 2008
     Darwin 1872, p. 421.
225. Edwards, A. W. F. (1 April 2000). "The Genetical Theory of Natural Selection". Genetics. 154 (4): 1419–1426. doi:10.1093/genetics/154.4.1419. PMC 1461012. PMID 10747041. Archived from the original on 24 September 2015.
226. FitzRoy 1839, pp. 216–218.
227. Leff 2000, Darwin's Timeline
228. "Territory origins". Northern Territory Department of Planning and Infrastructure, Australia. Archived from the original on 18 September 2006. Retrieved 15 December 2006.
229. Heard, Stephen B. (2020). Charles Darwin's barnacle and David Bowie's spider : how scientific names celebrate adventurers, heroes, and even a few scoundrels. Damstra, Emily S. New Haven: Yale University Press. ISBN 978-0-300-25269-9. OCLC 1143645266.
230. "Charles Darwin 200 years – Things you didn't know about Charles Darwin". Archived from the original on 28 May 2009. Retrieved 23 May 2009.
231. "The Darwin-Wallace Medal". The Linnean Society. 1 February 2016. Archived from the original on 29 March 2019. Retrieved 22 April 2022.
232. "Darwin College – Maps and directions – University of Kent". www.kent.ac.uk. Archived from the original on 31 October 2016. Retrieved 30 October 2016.
233. "How to join the noteworthy". BBC News. 7 November 2000. Archived from the original on 30 June 2006. Retrieved 24 April 2022.
234. "Author Jane Austen to feature on new £10 note". CBBC Newsround. 24 July 2013. Archived from the original on 24 April 2022. Retrieved 24 April 2022.
235. "Bank of England – Banknotes – Current Banknotes – £10". bankofengland.co.uk. 25 May 2005. Archived from the original on 25 May 2005. Retrieved 24 April 2022.
236. Darwin 1859, p. 434
237. Darwin 1859, p. 479
238. Darwin 1871, p. 1
239. Darwin 1874, p. vi
240. Darwin 1871, pp. 214, 232

Bibliography

• Anonymous (1882). "Obituary: Death Of Chas. Darwin". The New York Times. No. 21 April 1882. Archived from the original on 15 October 2009. Retrieved 30 October 2008.
• Balfour, J. H. (11 May 1882). "Obituary Notice of Charles Robert Darwin" . Transactions & Proceedings of the Botanical Society of Edinburgh (14): 284–298.
• Bannister, Robert C. (1989). Social Darwinism: Science and Myth in Anglo-American Social Thought. Philadelphia: Temple University Press. ISBN 978-0-87722-566-9.
• Bowler, Peter J. (2003). Evolution: The History of an Idea (3rd ed.). University of California Press. ISBN 978-0-520-23693-6.
• Browne, E. Janet (1995). Charles Darwin: vol. 1 Voyaging. London: Jonathan Cape. ISBN 978-1-84413-314-7.
• Browne, E. Janet (2002). Charles Darwin: vol. 2 The Power of Place. London: Jonathan Cape. ISBN 978-0-7126-6837-8.
• Darwin, Charles (1 December 1835). Henslow, J S (ed.). [Extracts from letters addressed to Professor Henslow]. Cambridge: . Archived from the original on 4 October 2022. Retrieved 27 January 2022.
• Darwin, Charles (1837). Notebook B: (Transmutation of species). Darwin Online. CUL-DAR121. Archived from the original on 8 February 2009. Retrieved 20 December 2008.
• Darwin, Charles (1839). Narrative of the surveying voyages of His Majesty's Ships Adventure and Beagle between the years 1826 and 1836, describing their examination of the southern shores of South America, and the Beagle's circumnavigation of the globe. Journal and remarks. 1832–1836. Vol. III. London: Henry Colburn. Archived from the original on 15 August 2012. Retrieved 24 October 2008.
• Darwin, Charles (1842). "Pencil Sketch of 1842". In Darwin, Francis (ed.). The foundations of The origin of species: Two essays written in 1842 and 1844. Cambridge University Press (published 1909). ISBN 978-0-548-79998-7. Archived from the original on 29 September 2011. Retrieved 13 December 2006.
• Darwin, Charles (1845). Journal of researches into the natural history and geology of the countries visited during the voyage of H.M.S. Beagle round the world, under the Command of Capt. Fitz Roy, R.N. 2d edition. London: John Murray. Archived from the original on 17 September 2011. Retrieved 24 October 2008.
• Darwin, Charles (1859). On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life (1st ed.). London: John Murray. ISBN 978-1-4353-9386-8. Archived from the original on 5 October 2008. Retrieved 24 October 2008.
• Darwin, Charles (1871). The Descent of Man, and Selection in Relation to Sex (1st ed.). London: John Murray. ISBN 978-0-8014-2085-6. Archived from the original on 12 July 2011. Retrieved 24 October 2008.
• Darwin, Charles (1872). The Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life (6th ed.). London: John Murray. ISBN 978-1-4353-9386-8. Archived from the original on 7 January 2010. Retrieved 1 November 2009.
• Darwin, Charles (1874). The Descent of Man, and Selection in Relation to Sex (2nd ed.). London: John Murray. ISBN 978-0-8014-2085-6. Archived from the original on 12 July 2011. Retrieved 16 January 2016.
• Darwin, Charles (1887). Darwin, Francis (ed.). The life and letters of Charles Darwin, including an autobiographical chapter. London: John Murray. ISBN 978-0-404-08417-2. Archived from the original on 5 March 2011. Retrieved 4 November 2008.
• Darwin, Charles (1958). Barlow, Nora (ed.). The Autobiography of Charles Darwin 1809–1882. With the original omissions restored. Edited and with appendix and notes by his granddaughter Nora Barlow. London: Collins. Archived from the original on 16 August 2013. Retrieved 28 September 2013.
• Darwin, Charles (2006). "Journal". In van Wyhe, John (ed.). Darwin's personal 'Journal' (1809–1881). Darwin Online. CUL-DAR158.1–76. Archived from the original on 24 December 2008. Retrieved 20 December 2008.
• Darwin, Charles; Costa, James T. (2009). The Annotated Origin: A Facsimile of the First Edition of On the Origin of Species Annotated by James T. Costa. Cambridge, Massachusetts, and London, England: Belknap Press of Harvard University Press. ISBN 978-0-674-03281-1.
• Desmond, Adrian; Moore, James (1991). Darwin. London: Michael Joseph, Penguin Group. ISBN 978-0-7181-3430-3.
• Desmond, Adrian; Moore, James; Browne, Janet (2004). "Darwin, Charles Robert". Oxford Dictionary of National Biography (online ed.). Oxford, England: Oxford University Press. doi:10.1093/ref:odnb/7176. ISBN 978-0-19-861411-1. (Subscription or UK public library membership required.)
• Desmond, Adrian; Moore, James; Browne, Janet (2007). Charles Darwin (Very Interesting People). Oxford, England: Oxford University Press. ISBN 978-0-19-921354-2.
• Desmond, Adrian; Moore, James (2009). Darwin's sacred cause : race, slavery and the quest for human origins. London: Allen Lane. ISBN 978-1-84614-035-8.
• Eldredge, Niles (2006). "Confessions of a Darwinist". The Virginia Quarterly Review (Spring 2006): 32–53. Archived from the original on 24 December 2013. Retrieved 4 November 2008.
• FitzRoy, Robert (1839). Voyages of the Adventure and Beagle, Volume II. London: Henry Colburn. Archived from the original on 5 May 2011. Retrieved 4 November 2008.
• Freeman, R. B. (1977). The Works of Charles Darwin: An Annotated Bibliographical Handlist. Folkestone: Wm Dawson & Sons Ltd. ISBN 978-0-208-01658-4. Archived from the original on 4 October 2022. Retrieved 4 November 2008.
• Freeman, R. B. (2007). Charles Darwin: A companion (2nd online ed.). The Complete Works of Charles Darwin Online. pp. 107, 109. Archived from the original on 25 December 2014. Retrieved 25 December 2014.
• Herbert, Sandra (1980). "The red notebook of Charles Darwin". Bulletin of the British Museum (Natural History), Historical Series. 7 (7 (24 April)): 1–164. doi:10.5962/p.272299. Archived from the original on 11 July 2007. Retrieved 11 January 2009.
• Herbert, Sandra (1991). "Charles Darwin as a prospective geological author". British Journal for the History of Science. 24 (2): 159–192. doi:10.1017/S0007087400027060. S2CID 143748414. Archived from the original on 29 March 2017. Retrieved 24 October 2008.
• Holliday, Trenton (2021). Cro-Magnon: The Story of the Last Ice Age People of Europe. NewYork: Columbia University Press. ISBN 978-0-231-20497-2.
• Huxley, Julian; Kettlewell, H.B.D. (1965). Charles Darwin and His World. New York: the Viking Press.
• Keynes, Richard (2000). Charles Darwin's zoology notes & specimen lists from H.M.S. Beagle. Cambridge University Press. ISBN 978-0-521-46569-4. Archived from the original on 5 December 2008. Retrieved 22 November 2008.
• Keynes, Richard (2001). Charles Darwin's Beagle Diary. Cambridge University Press. ISBN 978-0-521-23503-7. Archived from the original on 4 June 2012. Retrieved 24 October 2008.
• Kotzin, Daniel (2004). "Point-Counterpoint: Social Darwinism". Columbia American History Online. Archived from the original on 19 July 2011. Retrieved 22 November 2008.
• Larson, Edward J. (2004). Evolution: The Remarkable History of a Scientific Theory. Modern Library. ISBN 978-0-679-64288-6.
• Leff, David (2000). "AboutDarwin.com" (2000–2008 ed.). Archived from the original on 28 August 2013. Retrieved 30 December 2008.
• Leifchild (19 November 1859). "Review of 'Origin'". Athenaeum (1673). Archived from the original on 5 December 2008. Retrieved 22 November 2008.
• Miles, Sara Joan (2001). "Charles Darwin and Asa Gray Discuss Teleology and Design". Perspectives on Science and Christian Faith. 53: 196–201. Archived from the original on 5 April 2020. Retrieved 22 November 2008.
• Moore, James (2005). "Darwin – A 'Devil's Chaplain'?" (PDF). American Public Media. Archived from the original (PDF) on 27 February 2008. Retrieved 22 November 2008.
• Moore, James (2006). "Evolution and Wonder – Understanding Charles Darwin". Speaking of Faith (Radio Program). American Public Media. Archived from the original on 22 December 2008. Retrieved 22 November 2008.
• Owen, Richard (1840). Darwin, C. R. (ed.). Fossil Mammalia Part 1. The zoology of the voyage of H.M.S. Beagle. London: Smith Elder and Co.
• Paul, Diane B. (2003). "Darwin, social Darwinism and eugenics". In Hodge, Jonathan; Radick, Gregory (eds.). The Cambridge Companion to Darwin. Cambridge University Press. pp. 214–239. ISBN 978-0-521-77730-8.
• Radick, Gregory (2013). "Darwin and Humans". In Ruse, Michael (ed.). The Cambridge Encyclopedia of Darwin and Evolutionary Thought. Cambridge University Press. pp. 173–181.
• Smith, Charles H. (1999). "Alfred Russel Wallace on Spiritualism, Man, and Evolution: An Analytical Essay". Archived from the original on 5 December 2008. Retrieved 7 December 2008.
• Sulloway, Frank J. (1982). "Darwin and His Finches: The Evolution of a Legend" (PDF). Journal of the History of Biology. 15 (1): 1–53. CiteSeerX 10.1.1.458.3975. doi:10.1007/BF00132004. S2CID 17161535. Archived from the original (PDF) on 16 December 2008. Retrieved 9 December 2008.
• Sweet, William (2004). "Herbert Spencer". Internet Encyclopedia of Philosophy. Archived from the original on 28 May 2010. Retrieved 16 December 2008.
• Wilkins, John S. (1997). "Evolution and Philosophy: Does evolution make might right?". TalkOrigins Archive. Archived from the original on 14 May 2011. Retrieved 22 November 2008.
• Wilkins, John S. (2008). "Darwin". In Tucker, Aviezer (ed.). A Companion to the Philosophy of History and Historiography. Blackwell Companions to Philosophy. Chichester: Wiley-Blackwell. pp. 405–415. ISBN 978-1-4051-4908-2.
• van Wyhe, John (27 March 2007). "Mind the gap: Did Darwin avoid publishing his theory for many years?". Notes and Records of the Royal Society. 61 (2): 177–205. doi:10.1098/rsnr.2006.0171. S2CID 202574857. Archived from the original on 11 January 2011. Retrieved 7 February 2008.
• van Wyhe, John (2008). "Charles Darwin: gentleman naturalist: A biographical sketch". Darwin Online. Archived from the original on 13 January 2020. Retrieved 17 November 2008.
• van Wyhe, John (2008b). Darwin: The Story of the Man and His Theories of Evolution. London: Andre Deutsch Ltd (published 1 September 2008). ISBN 978-0-233-00251-4.
• von Sydow, Momme (2005). "Darwin – A Christian Undermining Christianity? On Self-Undermining Dynamics of Ideas Between Belief and Science" (PDF). In Knight, David M.; Eddy, Matthew D. (eds.). Science and Beliefs: From Natural Philosophy to Natural Science, 1700–1900. Burlington: Ashgate. pp. 141–156. ISBN 978-0-7546-3996-1. Archived from the original (PDF) on 26 March 2009. Retrieved 16 December 2008.
• Yates, Simon (2003). "The Lady Hope Story: A Widespread Falsehood". TalkOrigins Archive. Archived from the original on 12 October 2009. Retrieved 15 December 2006.

External links

Library resources about
Charles Darwin

---

• Online books
• Resources in your library
• Resources in other libraries

By Charles Darwin

• Online books
• Resources in your library
• Resources in other libraries

• "The Complete Work of Charles Darwin Online". Retrieved 4 March 2024.
• Works by Charles Darwin in eBook form at Standard Ebooks
• Works by Charles Darwin at Project Gutenberg
• Works by or about Charles Robert Darwin at the Internet Archive
• Works by Charles Darwin at LibriVox (public domain audiobooks)
• The Complete Works of Charles Darwin Online – Darwin Online; Darwin's publications, private papers and bibliography, supplementary works including biographies, obituaries and reviews
• Darwin Correspondence Project Full text and notes for complete correspondence to 1867, with summaries of all the rest, and pages of commentary
• Darwin Manuscript Project
• "Archival material relating to Charles Darwin". UK National Archives.
• View books owned and annotated by Charles Darwin at the online Biodiversity Heritage Library.
• Works by Charles Darwin at the Biodiversity Heritage Library
• Digitised Darwin Manuscripts in Cambridge Digital Library
• Portraits of Charles Darwin at the National Portrait Gallery, London
• Newspaper clippings about Charles Darwin in the 20th Century Press Archives of the ZBW
• Charles Darwin in the British horticultural press – Occasional Papers from RHS Lindley Library, volume 3 July 2010
• Scientific American, 29 April 1882, pp. 256, Obituary of Charles Darwin
• "Charles Darwin". Internet Encyclopedia of Philosophy.

v t e Charles Darwin
Life	Darwin–Wedgwood family Erasmus Darwin Josiah Wedgwood Education Voyage on HMS Beagle Inception of theory Development of theory Publication of theory Reactions to On the Origin of Species Orchids to Variation Descent of Man to Emotions Insectivorous Plants to Worms Religious views Health Women Portraits of Darwin (caricatures)
Writings	Extracts from Letters to Henslow (1835) The Voyage of the Beagle (1839) The Structure and Distribution of Coral Reefs (1842) Zoology of the Voyage of H.M.S. Beagle (1838–1843) Geological Observations on the Volcanic Islands (1844) Geological Observations on South America (1846) "On the Tendency of Species to form Varieties" (1858) On the Origin of Species (1859) Fertilisation of Orchids (1862) The Variation of Animals and Plants Under Domestication (1868) Natural Selection The Descent of Man, and Selection in Relation to Sex (1871) The Expression of the Emotions in Man and Animals (1872) Insectivorous Plants (1875) On the Movements and Habits of Climbing Plants (1875) The Effects of Cross and Self Fertilisation in the Vegetable Kingdom (1876) The Different Forms of Flowers on Plants of the Same Species (1877) The Power of Movement in Plants (1880) The Formation of Vegetable Mould Through the Action of Worms (1881) The Autobiography of Charles Darwin (1887) Correspondence The Life and Letters of Charles Darwin, 1887 More Letters of Charles Darwin, 1903 List of described taxa
Related	Eugenics Dysgenics Vestiges of the Natural History of Creation The Genetical Theory of Natural Selection (1930) History of evolutionary thought Pangenesis Darwin Industry Commemoration things named for Darwin Darwinism Neural Quantum Social Universal Alternatives to Eclipse of HMS Beagle Darwin Medal Darwin Awards Huxley family

Links to related articles

v t e Evolutionary biology Introduction Outline Timeline of evolution History of life Index Evolution Abiogenesis Adaptation Adaptive radiation Altruism Cheating Reciprocal Baldwin effect Cladistics Coevolution Mutualism Common descent Convergence Divergence Earliest known life forms Evidence of evolution Evolutionary arms race Evolutionary pressure Exaptation Extinction Event Homology Last universal common ancestor Macroevolution Microevolution Mismatch Non-adaptive radiation Origin of life Panspermia Parallel evolution Signalling theory Handicap principle Speciation Species Species complex Taxonomy Unit of selection Gene-centered view of evolution Population genetics Artificial selection Biodiversity Evolutionarily stable strategy Fisher's principle Fitness Inclusive Gene flow Genetic drift Kin selection Parental investment Parent–offspring conflict Mutation Population Natural selection Sexual dimorphism Sexual selection Flowering plants Fungi Mate choice Social selection Trivers–Willard hypothesis Variation Development Canalisation Evolutionary developmental biology Genetic assimilation Inversion Modularity Phenotypic plasticity Of taxa Bacteria Birds origin Brachiopods Molluscs Cephalopods Dinosaurs Fish Fungi Insects butterflies Life Mammals cats canids wolves dogs hyenas dolphins and whales horses Kangaroos primates humans lemurs sea cows Plants pollinator-mediated Reptiles Spiders Tetrapods Viruses Of organs Cell DNA Flagella Eukaryotes symbiogenesis chromosome endomembrane system mitochondria nucleus plastids In animals eye hair auditory ossicle nervous system brain Of processes Aging Death Programmed cell death Avian flight Biological complexity Cooperation Color vision in primates Emotion Empathy Ethics Eusociality Immune system Metabolism Monogamy Morality Mosaic evolution Multicellularity Sexual reproduction Gamete differentiation/sexes Life cycles/nuclear phases Mating types Meiosis Sex-determination Snake venom Tempo and modes Gradualism/Punctuated equilibrium/Saltationism Micromutation/Macromutation Uniformitarianism/Catastrophism Speciation Allopatric Anagenesis Catagenesis Cladogenesis Cospeciation Ecological Hybrid Non-ecological Parapatric Peripatric Reinforcement Sympatric History Renaissance and Enlightenment Transmutation of species David Hume Dialogues Concerning Natural Religion Charles Darwin On the Origin of Species History of paleontology Transitional fossil Blending inheritance Mendelian inheritance The eclipse of Darwinism Neo-Darwinism Modern synthesis History of molecular evolution Extended evolutionary synthesis Philosophy Darwinism Alternatives Catastrophism Lamarckism Orthogenesis Mutationism Saltationism Structuralism Spandrel Theistic Vitalism Teleology in biology Related Biogeography Ecological genetics Evolutionary medicine Group selection Cultural evolution Cultural group selection Dual inheritance theory Hologenome theory of evolution Missing heritability problem Molecular evolution Astrobiology Phylogenetics Tree Polymorphism Protocell Systematics Transgenerational epigenetic inheritance Category Portal v t e Ethology Branches Animal cognition Animal communication Animal consciousness Animal culture Animal sexual behaviour Animal welfare science Anthrozoology Bee learning and communication Behavioural ecology Behavioral endocrinology Behavioural genetics Breed Cognitive ethology Comfort behaviour Grooming Comparative psychology Emotion in animals Ethogram Evolutionary neuroscience Feeding Hover Human ethology Instinct Learning Neuroethology Pain in animals Philosophical ethology Sociobiology Stereotypy Structures Hive Honeycomb Nest Instinct Swarm Tool use by non-humans Zoosemiotics Zoomusicology Ethologists Patrick Bateson Marc Bekoff Donald Broom John B. Calhoun Charles Darwin Marian Dawkins Richard Dawkins Irenäus Eibl-Eibesfeldt Dian Fossey Karl von Frisch Jane Goodall Heini Hediger Julian Huxley Konrad Lorenz Desmond Morris Thomas Sebeok William Homan Thorpe Nikolaas Tinbergen Jakob von Uexküll Wolfgang Wickler E. O. Wilson Solly Zuckerman Societies Association for the Study of Animal Behaviour International Society for Applied Ethology Journals Animal Behaviour Animal Cognition Animal Welfare Behavioral Ecology Behaviour Category v t e Zoology Outline Branches Anthrozoology Arthropodology (Arachnology (Acarology) Carcinology Entomology (Coleopterology Lepidopterology Melittology Myrmecology Orthopterology) Myriapodology) Biological anthropology Bryozoology Cnidariology Ethnozoology Ethology Helminthology Herpetology (Batrachology Ophiology Testudinology) Ichthyology Malacology (Conchology Teuthology) Mammalogy (Cetology Cynology Felinology Hippology Primatology) Nematology Neuroethology Ornithology Parasitology Paleozoology Planktology Zooarchaeology Animal groups Porifera Ctenophora Placozoa Cnidaria Xenacoelomorpha Ambulacraria Chordata Aschelminthes Arthropoda Platyhelminthes Mollusca Annelida Animal anatomy Animal morphology Histology Tissues Epithelial tissue Muscular tissue Nervous tissue Connective tissues Mineralized tissues Molecular anatomy Anatomy and morphology Mammalian anatomy and morphology Human Dog Elephant Cat Bird anatomy Fish anatomy Shark anatomy Spider anatomy Insect morphology Animal physiology General physiology Respiration Respiratory system Breathing Gas exchange Respiratory pigment Cellular respiration Vascular system Blood Lymph Blood vessels Arteries Veins Capillaries Heart Water vascular system By species Insect physiology Fish physiology Zoologists Karl Ernst von Baer Georges Cuvier Charles Darwin Jean-Henri Fabre William Kirby Carl Linnaeus Konrad Lorenz Thomas Say Jakob von Uexküll Alfred Russel Wallace History Pre-Darwin Post-Darwin Timeline of zoology Category v t e Evolutionary psychologists Evolutionary psychology Biologists / neuroscientists Bernard Crespi John Crook Charles Darwin Richard Dawkins Jared Diamond W. D. Hamilton Alfred Kinsey Peter Kropotkin Gordon Orians Jaak Panksepp Margie Profet Peter Richerson Giacomo Rizzolatti Randy Thornhill Robert Trivers Carel van Schaik Claus Wedekind Mary Jane West-Eberhard Wolfgang Wickler George C. Williams David Sloan Wilson E. O. Wilson Richard Wrangham Anthropologists Jerome H. Barkow Christopher Boehm Robert Boyd Donald E. Brown Napoleon Chagnon Robin Dunbar Daniel Fessler Mark Flinn John D. Hawks Joseph Henrich Ruth Mace Daniel Nettle Stephen Shennan Donald Symons John Tooby Pierre van den Berghe Psychologists / cognitive scientists Mary Ainsworth Simon Baron-Cohen Justin L. Barrett Jay Belsky Jesse Bering David F. Bjorklund Paul Bloom John Bowlby Pascal Boyer Joseph Bulbulia David Buss Josep Call Anne Campbell Donald T. Campbell Peter Carruthers Noam Chomsky Leda Cosmides Martin Daly Paul Ekman Bruce J. Ellis Anne Fernald Aurelio José Figueredo Diana Fleischman Uta Frith Gordon G. Gallup David C. Geary Gerd Gigerenzer Peter Gray Jonathan Haidt Harry Harlow Judith Rich Harris Martie Haselton Stephen Kaplan Douglas T. Kenrick Simon M. Kirby Robert Kurzban Brian MacWhinney Michael T. McGuire Geoffrey Miller Darcia Narvaez Katherine Nelson Randolph M. Nesse Steven Neuberg David Perrett Henry Plotkin Steven Pinker Paul Rozin Mark Schaller David P. Schmitt Nancy Segal Todd K. Shackelford Roger Shepard Irwin Silverman Peter K. Smith Dan Sperber Anthony Stevens Frank Sulloway Michael Tomasello Joshua Tybur Mark van Vugt Andrew Whiten Glenn Wilson Margo Wilson Other social scientists Christopher Badcock Samuel Bowles Ernst Fehr Herbert Gintis Dominic D. P. Johnson Gad Saad Literary theorists / philosophers Edmund Burke Joseph Carroll Daniel Dennett Denis Dutton Thomas Hobbes David Hume Research centers/ organizations Center for Evolutionary Psychology Human Behavior and Evolution Society Max Planck Institute for Evolutionary Anthropology Max Planck Institute for Human Cognitive and Brain Sciences New England Complex Systems Institute Publications The Adapted Mind Evolution and Human Behavior The Evolution of Human Sexuality Evolution, Mind and Behaviour Evolutionary Behavioral Sciences Evolutionary Psychology  Evolutionary psychology  Psychology portal  Evolutionary biology portal v t e Copley Medallists (1851–1900) Richard Owen (1851) Alexander von Humboldt (1852) Heinrich Wilhelm Dove (1853) Johannes Peter Müller (1854) Léon Foucault (1855) Henri Milne-Edwards (1856) Michel Eugène Chevreul (1857) Charles Lyell (1858) Wilhelm Eduard Weber (1859) Robert Bunsen (1860) Louis Agassiz (1861) Thomas Graham (1862) Adam Sedgwick (1863) Charles Darwin (1864) Michel Chasles (1865) Julius Plücker (1866) Karl Ernst von Baer (1867) Charles Wheatstone (1868) Henri Victor Regnault (1869) James Prescott Joule (1870) Julius Robert von Mayer (1871) Friedrich Wöhler (1872) Hermann von Helmholtz (1873) Louis Pasteur (1874) August Wilhelm von Hofmann (1875) Claude Bernard (1876) James Dwight Dana (1877) Jean-Baptiste Boussingault (1878) Rudolf Clausius (1879) James Joseph Sylvester (1880) Charles Adolphe Wurtz (1881) Arthur Cayley (1882) William Thomson (1883) Carl Ludwig (1884) Friedrich August Kekulé von Stradonitz (1885) Franz Ernst Neumann (1886) Joseph Dalton Hooker (1887) Thomas Henry Huxley (1888) George Salmon (1889) Simon Newcomb (1890) Stanislao Cannizzaro (1891) Rudolf Virchow (1892) George Gabriel Stokes (1893) Edward Frankland (1894) Karl Weierstrass (1895) Carl Gegenbaur (1896) Albert von Kölliker (1897) William Huggins (1898) John William Strutt (1899) Marcellin Berthelot (1900) v t e Fellows of the Royal Society elected in 1839 Fellows Thomas Dyke Acland George Barker Beriah Botfield Robert Carrington Arthur Conolly Charles Darwin Edward Davies Davenport Henry Mangles Denham Richard Drew Henry Drummond Arthur Farre Thomas William Fletcher William James Frodsham Thomas Gaskin George Godwin John T. Graves Edwin Guest George Gulliver James Halliwell-Phillipps Peter Hardy James Heywood John Hilton John Hogg Gilbert Wakefield Mackmurdo Samuel Roffey Maitland Henry Moseley H. Alexander Ormsby William Reid Robert Rigg John Rogers George Leith Roupell William Sharpey Clement Tudway Swanston James Joseph Sylvester Charles Thorp Charles Turnor John Wesley Williams James Yates Foreign Christopher Hansteen Macedonio Melloni Adolphe Quetelet Félix Savart v t e Nonverbal communication Modalities Physical Blushing Body language / Kinesics Body-to-body communication Facial expression Facial Action Coding System Microexpression Subtle expression Gesture List Speech-independent gestures Haptic communication Imitation Interpersonal synchrony Laughter Oculesics Eye contact Pupil dilation Olfaction Posture Proxemics Speech Affect Emotional prosody Paralanguage Intonation Loudness Prosody Rhythm Stress Tone Voice quality Social context Chronemics Conventions Display rules Habitus High-context and low-context cultures Interpersonal relationship Social norm Other Emoticon / Smiley One-bit message Missed call Silent service code Unconscious Microexpression Non-verbal leakage Multi-faceted Affect display Deception Emotion recognition First impression Intimacy Broader concepts Basic interpersonal communicative skills Communication Emotional intelligence Nunchi People skills Semiotics Social behavior Social cue Social competence Social skills Unsaid Further information Disorders Aprosodia Asperger syndrome Autism Fragile X Pervasive developmental disorder not otherwise specified Childhood disintegrative disorder Rett syndrome Dyssemia Nonverbal learning disorder Social (pragmatic) communication disorder Neuroanatomy Limbic system / Limbic lobe Mirror neuron Applications Cold reading Lie detection Freudian slip Poker tell Targeted advertising Technology Computer processing of body language Emotion recognition in conversation Gesture recognition List of facial expression databases Sentiment analysis Key people Ray Birdwhistell Charles Darwin Paul Ekman Related Animal communication Behavioral communication Aggressive Assertive Passive Passive-aggressive Impression management Meta-communication Monastic sign lexicons Verbal communication Non-verbal language Sign language Tactile signing Tadoma Art and literature Mime Mimoplastic art Subtext

Charles Darwin at Misplaced Pages's sister projects:

• Definitions from Wiktionary
• Media from Commons
• Quotations from Wikiquote
• Texts from Wikisource
• Textbooks from Wikibooks
• Taxa from Wikispecies
• Data from Wikidata

Categories:

• Charles Darwin
• 1809 births
• 1882 deaths
• 19th-century British biologists
• 19th-century English writers
• 19th-century Anglicans
• 19th-century English naturalists
• 19th-century British geologists
• Alumni of Christ's College, Cambridge
• Alumni of the University of Edinburgh
• British carcinologists
• Burials at Westminster Abbey
• Circumnavigators of the globe
• Coleopterists
• Darwin–Wedgwood family
• Deaths from coronary thrombosis
• English abolitionists
• English agnostics
• English Anglicans
• English entomologists
• English geologists
• English justices of the peace
• English sceptics
• English travel writers
• Ethologists
• British evolutionary biologists
• Fellows of the Linnean Society of London
• Fellows of the Royal Entomological Society
• Fellows of the Royal Geographical Society
• Fellows of the Royal Society
• Fellows of the Zoological Society of London
• Human evolution theorists
• Human evolution
• Independent scientists
• Members of the American Philosophical Society
• Members of the Lincean Academy
• Members of the Royal Academy of Belgium
• Members of the Royal Netherlands Academy of Arts and Sciences
• Members of the Royal Swedish Academy of Sciences
• People educated at Shrewsbury School
• Scientists from Shrewsbury
• Recipients of the Copley Medal
• Recipients of the Pour le Mérite (civil class)
• Royal Medal winners
• Theoretical biologists
• Wollaston Medal winners